Interesting Facts
- This species is famous for obtaining undischarged cnidae (cells which bear nematocysts) from its Cnidarian prey and moving them through the hepatic diverticula to the tips of the cerata , where they are likely used for defense. If disturbed they sometimes wave their cerata. If one of the cerata is broken off, muscles within it contract, expelling the nematocysts, which then discharge .
Common Names
Common Names in English:
Common Grey Sea Slug, Maned Nudibranch, Mossy Nudibranch, Papillose Aeolid, Sea Mouse, Shag Rug Nudibranch, Shag-Rug Aeolis, Shaggy Mouse Nudibranch
Description
Physical Description
Species Aeolidia papillosa
Like other Aeolid nudibranchs, this species has its anus on the right side of the body on an inconspicuous tubercle, it has dorsal outgrowths such as cerata (which are cylindrical, lanceolate, or leaflike but not bushy and are usually arranged in transverse rows ), the rhinophores stand free and cannot be retracted into a sheath . Aeolidia papillosa has many flattened, lanceolate cerata but a bare area down the middle of the dorsum with no cerata. It has no sail-like ridge on the posterior side of its cerata, at least several rows of cerata are based anterior to the rhinophores, the cerata are less than 1/3 the length of the body, the tail tapers to a blunt point (is not sharply drawn out into a point) (photo), the anterolateral corners of the foot project as pedal tentacles (photo). The body and cerata are covered with gray to brown spots, especially on the cerata (some animals may lack pigment). Usually there is a bright white triangular patch on the dorsum over the head just anterior to the rhinophores (photo). The rhinophores are smooth and taper gradually from the base to the tip , which appears to have a pore in it. Although hepatic diverticula from the digestive system extend into each of the cerata, the paths of these diverticula are not readily visible from outside if the animal is pigmented . Color variable but often light brownish, brownish-cream, white, gray, or pink with lighter spots down the center of the dorsum. It often takes on some of the color of the anemones it is eating. Often 3-5 cm long, may attain 6 or even 10 cm.
Habitat
On rocks, or may be on floats or docks. Often near its perferred prey , Anthopleura elegantissima. Intertidal to 900 m
Typically found in the intertidal zone at the water's edge at a mean distance from sea level of 116 meters (381 feet).[1]
Biology
Diet
Feeds on anemones, especially Anthopleuraelegantissima and secondarily Metridium senile. Also may feed on Urticina crassicornis, Anthopleura xanthogrammica, and Epiactis prolifera, the young of which it may swallow whole, as well as sea pens and hydroids . It can detect its prey from a distance . It apparently does not prey on Anthopleura artemisia. It is said to be a voracious predator , consuming enough anemone tissue to equal half or all its body weight per day. It preys on large anemones by first spreading mucus on the column, then biting off and swallowing chunks . The mucus may shield the nudibranch from nematocyst discharge , plus this species' mucus seems to elicit less nematocyst discharge than does the mucus from other, non-anemone-eating nudibranchs such as Hermissenda crassicornis or Cadlina luteomarginata so it may have some inhibitory effect. (Anemones may eat Hermissenda or Cadlina, but Aeolidia eats the anemone). Tough cuticle in the mouth and esophagus may protect those areas from nematocysts. It may eventually eat entire large anemones. After eating Anthopleura elegantissima which is symbiotic with algae, the algae may also be segregated into the tips of its cerata where they continue photosynthesis.
Reproduction
In SE Alaska this species reproduces late March to late April. It lays a white to pinkish, coiled string of eggs in capsules which are attached to rocks or eelgrass leaves. In Washington, eggs hatch as veligers after 10-24 days.
Taxonomy
- Domain:
Eukaryota
(
)
- Whittaker & Margulis,1978
- eukaryotes
- Kingdom:
Animalia
(
)
- Linnaeus, 1758
- animals
- Subkingdom:
Bilateria
(
)
- (Hatschek, 1888) Cavalier-Smith, 1983
- Branch:
Protostomia
(
)
- Grobben, 1908
- Infrakingdom:
Lophotrochozoa
(
)
- Superphylum:
Eutrochozoa
(
)
- Phylum:
Mollusca
(
)
- (Linnaeus, 1758) Cuvier, 1795
- Molluscs
- Class:
Gastropoda
(
)
- Cuvier, 1795
- Snails and Slugs
- Subclass:
Orthogastropoda
(
)
- Ponder & Lindberg, 1996
- Superorder:
Heterobranchia
(
)
- J.e. Gray, 1840
- Order:
Opisthobranchia
(
)
- Milne-Edwards, 1848
- Suborder:
Nudibranchia
(
)
- Blainville, 1814
- Superfamily:
Aeolidioidea
(
)
- Gray, 1827
- Family:
Aeolidiidae
(
)
- Gray, 1827
- Genus:
Aeolidia
(
)
- (Linnaeus, 1761)
- Specific name:
papillosa
- (Linnaeus, 1761)
- Scientific name: - Aeolidia papillosa (Linnaeus, 1761)
- Specific name:
papillosa
- (Linnaeus, 1761)
- Genus:
Aeolidia
(
- Family:
Aeolidiidae
(
- Superfamily:
Aeolidioidea
(
- Suborder:
Nudibranchia
(
- Order:
Opisthobranchia
(
- Superorder:
Heterobranchia
(
- Subclass:
Orthogastropoda
(
- Class:
Gastropoda
(
- Phylum:
Mollusca
(
- Superphylum:
Eutrochozoa
(
- Infrakingdom:
Lophotrochozoa
(
- Branch:
Protostomia
(
- Subkingdom:
Bilateria
(
- Kingdom:
Animalia
(
Unambiguous Synonyms
- Aeolidia farinacea
Notes
Name Status: Accepted Name . Latest taxonomic scrutiny: 17-Oct-2001.
Similar Species
Flabellina salmonacea looks similar but no cerata are attached anterior to the rhinophores. A deep-water species, A. herculea, lives at depths greater than 500 m.
Members of the genus Aeolidia
There are approximately 2 species in this genus:
A. helicochorda · A. papillosa (Shaggy Mouse Nudibranch)
More Info
- Search for Pictures: images.google.com
- Search for Scholarly Articles: Google Scholar
- Search using Scientific Name and Vernacular Names: All the Web | AltaVista Canada | AltaVista | Excite | Google | HotBot | Lycos
- Search using Specialized Databases: GenBank | Medline | Scirus | CISTI/CAL | Agricola Periodicals | Agricola Books
Further Reading
- Archiv für Naturgeschichte. Berlin: Nicolai, 1835- GER url p. 165.
- Arkiv för zoologi / utgivet af K. Svenska vetenskaps-akademien. Stockholm: P.A. Norstedt and soner, 1903-1974. SWE url p. 104.
- Biological survey of the Mount Desert Region, conducted by William Procter... From the Mount Desert Island Biological Laboratory, Mount Desert Island, Maine. Philadelphia, The Wistar Institute of Anatomy and Biology, 1927-1946. ENG url p. 176, p. 210.
- Boletín de la Sociedad Española de Historia Natural. Madrid: Estab. tip. de Fortenet, 1901-1937. SPA url p. 238.
- Bulletin - United States National Museum. Washington: Smithsonian Institution Press, [etc.];1877-1971. ENG url p. 146.
- Bulletin biologique de la France et de la Belgique. Montreuil [etc.]Gauther-Villars [etc.] FRE url p. 301.
- Det Kongelige Norske videnskabers selskabs skrifter. Trondhjem: Aktietrykkeriet, -1969. DAN url p. 13.
- Fauna of New England. Boston, Printed for the [Boston] Society [of Natural History] from the Gurdon Saltonstall Fund, 1904-25. ENG url p. 162.
- Lehrbuch der Zoologie / von Richard Hertwig. Jena: G. Fischer, 1892. GER url p. 331.
- Memoirs of the California Academy of Sciences. San Francisco: The Academy, 1888- ENG url p. 376, p. 542.
- Proceedings of the Academy of Natural Sciences of Philadelphia. [Philadelphia, Academy of Natural Sciences of Philadelphia] ENG url p. 71, p. 74, p. 75.
- Revista do Museu Paulista. São Paulo: Museu Paulista, 1895- POR url p. 420.
- Special occasional publication / Department of Mollusks, Museum of Comparative Zoology, Harvard University. Cambridge, Mass., The Museum, 1973- ENG url p. 14, p. 17, p. 49, p. 55, p. 57, p. 6.
- Special scientific report. Seattle, National Marine Fisheries Service; for sale by the Supt. of Docs., U.S. Govt. Print. Off., 1949-1971. ENG url p. 16.
- The Biological bulletin. Woods Hole, Mass.: Marine Biological Laboratory, ENG url p. 111, p. 113, p. 113, p. 114, p. 115, p. 115, p. 117, p. 119, p. 120, p. 121, p. 122, p. 123, p. 125, p. 131, p. 138, p. 139, p. 140, p. 141, p. 142, p. 144, p. 145, p. 151, p. 152, p. 192, p. 211, p. 212, p. 214, p. 216, p. 217, p. 218, p. 220, p. 221, p. 223, p. 223, p. 224, p. 224, p. 226, p. 227, p. 228, p. 229, p. 230, p. 252, p. 283, p. 285, p. 319, p. 357, p. 358, p. 359, p. 375, p. 394, p. 395, p. 397, p. 404, p. 405, p. 406, p. 407, p. 439, p. 480, p. 548, p. 614, p. 70, p. 73, p. 80, p. 816, p. 820, p. 830.
- The Nautilus. [Melbourne, Fla., etc., American Malacologists, inc., etc.] ENG url p. 109, p. 130, p. 26, p. 37, p. 38, p. 77, p. 80, p. 83.
- The biology of marine animals. New York, Interscience Publishers[1960] ENG url p. 252.
- Tijdschrift der Nederlandsche Dierkundige Vereeniging. Leiden, Brill. DUT url p. 10.
- Waters, V. L., 1973. Food-preference of the nudibranchAeolidia papillosa, and the effect of the defenses of the prey on predation. Veliger 15: 174-192
- Zoologischer Jahresbericht. Berlin, R. Friedlander. GER url p. 16, p. 29, p. 72.
- Zoologisches Centralblatt. Leipzig: W. Engelmann, [1894-1902] GER url p. 811.
Notes
Contributors
- Bisby, F.A., Y.R. Roskov, M.A. Ruggiero, T.M. Orrell, L.E. Paglinawan, P.W. Brewer, N. Bailly, J. van Hertum, eds (2007). Species 2000 and ITIS Catalogue of Life: 2007 Annual Checklist. Species 2000: Reading, U.K.
- Global Biodiversity Information Facility. Accessed December 11, 2007. http://www.gbif.org Mediated distribution data from 19 providers.
Data Sources
Accessed through GBIF Data Portal December 11, 2007:
- Academy of Natural Sciences, Academy of Natural Sciences Malacology Database
- Academy of Natural Sciences, Academy of Natural Sciences OBIS Mollusc Database [Darwin Core]
- Institute of Marine and Coastal Sciences, Rutgers University, A comparison of benthic biodiversity in the North Sea, English Channel and Celtic Seas
- Institute of Marine and Coastal Sciences, Rutgers University, Academy of Natural Sciences OBIS Mollusc Database
- Institute of Marine and Coastal Sciences, Rutgers University, Atlantic Reference Centre
- Institute of Marine and Coastal Sciences, Rutgers University, Bay of Fundy Species List
- Institute of Marine and Coastal Sciences, Rutgers University, BioMar
- Institute of Marine and Coastal Sciences, Rutgers University, Biogeography Scheldt Estuary
- Institute of Marine and Coastal Sciences, Rutgers University, Histroical benthic dredge samples from the southern baltic and the North Sea
- Institute of Marine and Coastal Sciences, Rutgers University, Taxonomic Information Sytem for the Belgian coastal area
- The Swedish Museum of Natural History
- , Bugs
- UK National Biodiversity Network, Countryside Council for Wales - Benthic Marine data from CCW Technical Support
- Contracts
- UK National Biodiversity Network, Marine Biological Association - Marine Life Survey Data
- collated by MarLIN
- UK National Biodiversity Network, Marine Biological Association - Marine survey data
- held by MarLIN.
Identifiers
- Biodiversity Heritage Library NamebankID: 2689388
- Catalogue of Life Accepted Name Code: ITS-78728
- Global Biodiversity Information Facility Taxonkey: 13979142
- Integrated Taxonomic Information System (ITIS) Taxonomic Serial Number (TSN): 78728
- Zipcode Zoo Species Identifier: 511432
Footnotes
- Standard Deviation = 922.950 based on 385 observations. Terrestrial altitude and ocean depth information for each observation from British Oceanographic Data Centre. [back]
