Trees evergreen, crown usually spirelike to conic, sometimes flat to round topped in age. Bark initially thin, smooth, bearing resin blisters, in age furrowed and/or flaking in plates. Branches whorled, irregularinternodal branches occasionally produced by epicormic sprouting (growing from a dormantbud) ; short (spur) shoots absent; leaf scarsprominent, ± circular to broadly elliptic, flush with twig surface, slightly depressed, or slightly raised evenly all around. Budsovate or oblong, resinous or not, apexrounded or pointed. Leaves borne singly, persisting 5 or more years, spirally arranged but often proximally twisted so as to appear either 1-ranked (pointing up like toothbrush bristles) or 2-ranked, sessile, typically constricted and often twisted above the somewhat broadened base, sheath absent; leaves on vegetative branches flattened, frequently groovedadaxially, usually notched to rounded at apex; leaves on fertile branches sometimes appearing 4-sided, upright, sharp-pointed to rounded at apex; resin canals 2. Cones borne on year-old twigs.Pollen cones grouped, ovate or oblong-cylindric, leaving gall-like protuberances after falling, yellow to red, green, blue, or purple. Seed cones maturing in 1 season, erect, ovoid to oblong-cylindric or cylindric, not falling whole but scale by scale, coneaxis persisting as an erect "spike" on branch; scales shed individually, fan-shaped, lacking apophysis and umbo; bractsincluded to exserted.Seedswinged, the wing-seed juncture bearing resin sac; cotyledons 4--10. x =12.
Species ca. 42: widespread in north temperate regions, North America, Mexico, Central America, Eurasia (s to Himalayas, s China, and Taiwan), n Africa.
In Abies several traditionally accepted species have closely allied sibling species, e.g., A. balsamea -- A. fraseri, A. bifolia -- A. lasiocarpa, and A. magnifica -- A. procera. Other species may be more distinct morphologically, but many of these still appear to have evolved in geographic isolation without strong reproductive barriers developing. Thus, when distributions of species overlap, introgression between the taxa is the rule; this may make it difficult to assign certain individuals to a species. In the interests of nomenclaturalstability, I have accepted the taxa recognized by the U.S. Forest Service (E.L. Little Jr. 1979). This classification does not recognize varieties based on variations in bract characteristics but recognizes species that perhaps would be treated as varieties in other conifergenera. The only exceptions to this treatment are some necessary changes within A. concolor and A. lasiocarpa. Cases of introgression are discussed under the taxa involved. Some distinct or possibly distinct geographic populations deserve further study and may warrant future taxonomic recognition.
Most North American firs are major components of vegetation, especially in the boreal, Pacific Coastconiferous, and western montane coniferous forests, where they are important for watershedmanagement. They are cut for pulpwood and lumber and, largely from plantations, for Christmas trees. All our species, especially Abies concolor, and several exotics are grown---some more than others---as ornamentals. Firs provide cover, and their leaves are important as food, for various birds and mammals. Species of Abies frequently have a pleasant odor; their foliage has been used as a stuffing material for pillows. Most commercial products with "pine odors" are in fact scented with essential oils distilled from Abies foliage by Russian farmers. A similar oil could be derived from balsam fir in North America.[1]
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