Overview
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Vulnerable |
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Description
Family Pinaceae
Trees
or rarely shrubs
, evergreen
or deciduous, monoecious. Branchlets
often dimorphic
: long branchlets with clearly spirally arranged
, sometimes scalelike leaves; short branchlets often reduced to slow growing lateral
spurs bearing dense clusters
of leaves at apex. Leaves solitary or in bundles of (1 or) 2-5(-8) when basally subtended by a leaf sheath
; leaf blade
linear
or needlelike, not decurrent. Cones unisexual
. Pollen cones solitary or clustered, with numerous
spirally arranged microsporophylls
; microsporophyll with 2 microsporangia; pollen usually 2-saccate (nonsaccate in Cedrus, Larix, Pseudotsuga, and most species of Tsuga) . Seed cones erect
or pendulous, maturing in 1st, 2nd, or occasionally 3rd year, dehiscent
or occasionally indehiscent, with many spirally arranged ovulate
scales
and bracts; ovulate scales usually smaller than bracts at pollination, with 2 upright ovules adaxially, free
or only basally adnate
with bracts, maturing into seed scales. Seed scales appressed, woody or leathery, variable in shape
and size, with 2 seeds adaxially, persistent
or deciduous after cone maturity. Bracts free or adnate basally with seed scales, well developed or rudimentary
, exserted or included
. Seeds terminally winged
(except in some species of Pinus) . Cotyledons 2-18. Germination hypogeal or epigeal. 2n = 24* (almost always) .
Ten or eleven genera and ca.
235 species: N hemisphere; ten genera (two endemic) and 108 species (43 endemic, 24 introduced
) in China.
Species of the Pinaceae are among the most valuable and commercially important plants
in the world. Most species are trees, and are often excellent sources of lumber, wood
products, and resins; many are cultivated for afforestation
and as ornamentals
.[1]
Genus Abies
Trees
evergreen
, crown usually spirelike to conic, sometimes flat to round topped in age. Bark
initially thin, smooth
, bearing resin blisters, in age furrowed
and/or flaking
in plates
. Branches whorled
, irregular internodal
branches occasionally produced
by epicormic
sprouting (growing from a dormant
bud) ; short (spur) shoots
absent; leaf scars
prominent
, ± circular to broadly elliptic
, flush with twig
surface, slightly depressed
, or slightly raised evenly all around. Buds ovate
or oblong
, resinous
or not, apex rounded
or pointed
. Leaves borne singly, persisting 5 or more years, spirally arranged
but often proximally twisted so as to appear either 1-ranked (pointing up like toothbrush bristles
) or 2-ranked, sessile, typically constricted
and often twisted above the somewhat broadened base
, sheath
absent; leaves on vegetative
branches flattened, frequently grooved
adaxially, usually notched
to rounded at apex; leaves on fertile
branches sometimes appearing 4-sided, upright, sharp-pointed to rounded at apex; resin canals 2. Cones borne on year-old twigs
. Pollen cones grouped, ovate or oblong-cylindric, leaving gall-like protuberances
after falling, yellow to red, green, blue, or purple. Seed cones maturing in 1 season
, erect
, ovoid
to oblong-cylindric or cylindric
, not falling whole but scale by scale, cone axis persisting as an erect "spike" on branch
; scales
shed individually, fan-shaped, lacking apophysis and umbo
; bracts included
to exserted. Seeds winged
, the wing-seed juncture bearing resin sac; cotyledons 4--10. x
=12.
Species ca.
42: widespread in north temperate regions
, North America, Mexico, Central America, Eurasia
(s to Himalayas, s China, and Taiwan), n Africa.
In Abies several traditionally accepted species have closely allied sibling species
, e.g.
, A. balsamea -- A. fraseri, A. bifolia -- A. lasiocarpa, and A. magnifica -- A. procera. Other species may be more distinct
morphologically, but many of these still appear to have evolved in geographic isolation
without strong
reproductive barriers
developing. Thus, when distributions of species overlap, introgression between the taxa is the rule
; this may make it difficult to assign certain individuals to a species. In the interests of nomenclatural
stability
, I have accepted the taxa recognized by the U.S. Forest
Service (E.L. Little Jr. 1979). This classification does not recognize varieties based on variations
in bract characteristics but recognizes species that perhaps would be treated as varieties in other conifer genera. The only exceptions to this treatment are some necessary changes within A. concolor
and A. lasiocarpa. Cases of introgression are discussed under the taxa involved. Some distinct or possibly distinct geographic populations deserve further study and may warrant future taxonomic
recognition.
Most North American firs are major components
of vegetation, especially in the boreal, Pacific Coast coniferous
, and western montane
coniferous forests, where they are important for watershed
management
. They are cut
for pulpwood and lumber and, largely from plantations, for Christmas trees
. All our species, especially Abies concolor, and several exotics
are grown---some more than others---as ornamentals
. Firs provide cover
, and their leaves are important as food, for various birds and mammals. Species of Abies frequently have a pleasant odor; their foliage
has been used as a stuffing material
for pillows
. Most commercial
products with "pine odors" are in fact scented with essential oils distilled from Abies foliage by Russian farmers. A similar oil
could be derived from balsam fir in North America.[2]
Habitat
Ecology: Populations are scattered and confined to areas of dense montane forest .[3]
Taxonomy
- Domain:
Eukaryota
(
)
- Whittaker & Margulis,1978
- eukaryotes
- Kingdom:
Plantae
(
)
- Haeckel, 1866
- Plants
- Subkingdom:
Viridaeplantae
(
)
- Cavalier-Smith, 1981
- Phylum:
Tracheophyta
(
)
- Sinnott, 1935 Ex Cavalier-Smith, 1998
- Vascular Plants
- Subphylum:
Euphyllophytina
(
)
- Infraphylum:
Radiatopses
(
)
- Kenrick & Crane, 1997
- Class:
Coniferopsida
(
)
- Order:
Coniferales
(
)
- Order:
Coniferales
(
- Class:
Coniferopsida
(
- Infraphylum:
Radiatopses
(
- Subphylum:
Euphyllophytina
(
- Phylum:
Tracheophyta
(
- Subkingdom:
Viridaeplantae
(
- Kingdom:
Plantae
(
Similar Species
Members of the genus Abies
There are approximately 507 species in this genus. Here are just 100 of them:
A. ajanensis · A. alba (European Silver Fir) · A. alba-Hybride · A. alba f. pendula (European Silver Fir) · A. alba 'Green Spiral' (Green Spiral Contorted Silver Fir) · A. alba 'King's Dwarf' · A. alba 'Nana' · A. alba 'Schwarzwald' · A. alba var. calabrica · A. albertiana · A. alcockiana · A. alcoqueana · A. alvordensis · A. amabilis (Pacific Silver Fir) · A. americana · A. apollinis · A. araragi · A. arizonica · A. arizonica 'Compacta' · A. arnoldiana · A. arnoldiana 'Cyrille' · A. balsamea (Canada Balsam) · A. balsamea f. hudsonia (Hudsonia Dwarf Silver Fir) · A. balsamea fraseri (Balsam Fir) · A. balsamea 'Hudsonia' · A. balsamea Hudsonia Group · A. balsamea 'Jamie' · A. balsamea lasiocarpa (Subalpine Fir) · A. balsamea 'Le Feber' · A. balsamea 'Renswoude' · A. lasiocarpa var. lasiocarpa · A. balsamea 'Tyler Blue' · A. balsamea var. balsamea · A. balsamea var. nana (Dwarf Balsam Fir) · A. balsamea var. phanerolepis · A. balsamea var. phanerolepis 'Bear Swamp' · A. balsamea 'Verkade's Prostrate' · A. balsamea x sibirica (Balsam Fir) · A. balsamen · A. batavorum · A. beissneriana · A. beshanzuensis · A. bicolor · A. bifolia · A. borisii-regis (King Boris Fir) · A. borisii-regis 'Pendula' · A. bornemuelleriana · A. brachyphylla · A. brachytyla · A. bracteata (Bristle-Cone Fir) · A. brunoniana · A. candicans · A. carpatica · A. cedroides · A. cephalonica (Grecian Fir) · A. cephalonica 'Compacta' · A. cephalonica 'Greg's Broom' · A. cephalonica 'Meyer's Dwarf' (Greek Fir) · A. cephalonica var. cephalonica · A. chaneyi · A. chengii · A. chensiensis (Shensi Fir) · A. chensiensis chensiensis · A. chensiensis salouenensis · A. chensiensis salouensis · A. chensiensis subsp. salouenensis · A. chensiensis subsp. yulongxueshanensis · A. chensiensis var. salouensis · A. chensiensis yulongxueshanensis · A. chiloensis · A. chilrowensis · A. chinensis · A. cilicica (Cilica Fir) · A. cilicica cilicica · A. cilicica isaurica · A. clambrasiliana · A. clanbrassiliana · A. coahuilensis · A. coerulescens · A. communis · A. concolor (Rocky Mountain White Fir) · A. concoloroides · A. concolor 'Archer's Dwarf' · A. concolor 'Argentea Wattezii' · A. concolor 'Birthday Broom' · A. concolor 'Blue Cloak' · A. concolor 'Blue Sapphire' · A. concolor 'Blue Spreader' · A. concolor 'Candicans' (Colorado Fir) · A. concolor 'Fagerhult' · A. concolor 'Gables Weeping' (Gable's Weeping Colorado Fir) · A. concolor 'Glauca Compacta' · A. concolor 'Glauca Nana' · A. concolor 'Glenmore' · A. concolor 'Green Globe' (Colorado Fir) · A. concolor 'Hillier Broom' · A. concolor 'Hillier's Dwarf' · A. concolor 'Husky Pup' · A. concolor lasiocarpa · A. concolor lowiana
More Info
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Further Reading
- Farjon, A. 1994. Annotations to: Conservation status listing of conifers of the world dated 2 February 1994.
- Farjon, A., Page, C.N. and Schellevis, N. 1993. A preliminary world list of threatened conifer taxa. Biodiversity and Conservation 2: 304-326.
- Hilton-Taylor, C. (compiler). 2000. 2000 IUCN Red List of Threatened Species. IUCN, Gland, Switzerland and Cambridge, UK.
- National Environment Protection Bureau. 1987. The list of rare and endangered plants protected in China. Botanical Institute of Chinese Academy of Sciences, Academy Press, Beijing.
- Oldfield, S., Lusty, C. and MacKinven, A. (compilers) 1998. The World List of Threatened Trees. World Conservation Press, Cambridge, UK.
- Cheng Wan-chün, Fu Li-kuo, Law Yu-wu, Fu Shu-hsia, Wang Wen-tsai, Chu Cheng-de, Chao Chi-son & Chen Chia-jui. 1978. Pinaceae. In: Cheng Wan-chün & Fu Li-kuo, eds., Fl. Reipubl. Popularis Sin. 7: 32-281.
- Liu, T. S. 1971. A Monograph of the Genus Abies. Taipei.
- Matzenko, A.E. 1968. Conspectus generis Abies Mill. Novosti Sist. Vyssh. Rast. 5: 9--12.
Notes
Contributors
- Conifer Specialist Group 2000. In IUCN 2008. 2008 IUCN Red List of Threatened Species. IUCNRedList.org. Downloaded July 19, 2008.
Identifiers
- Biodiversity Heritage Library NamebankID: 3497228
- IUCN ID: 34117
- Zipcode Zoo Species Identifier: 3963539
Footnotes
- Liguo Fu, Nan Li, Thomas S. Elias & Robert R. Mill "Pinaceae". in Flora of China Vol. 4 Page 11. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org. [back]
- Richard S. Hunt "Abies". in Flora of North America Vol. 2. Oxford University Press. Online at EFloras.org. [back]
- Conifer Specialist Group 2000. In IUCN 2008. 2008 IUCN Red List of Threatened Species. IUCNRedList.org. Downloaded July 19, 2008. [back]
