Overview
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Vulnerable |
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Common Names
Common Names in English:
Mojave Sand Verbena, Mojave Sandverbena
Description
Family Nyctaginaceae
Herbs, shrubs
, trees
, or sometimes spiny
vines
. Leaves opposite, alternate, or whorled
; stipules absent; petiole
usually present, well defined; leaf blade
simple
, herbaceous or slightly fleshy
, margin
entire. Inflorescences mostly terminal
, less often axillary
, of cymes, umbels, or verticils
, sometimes 1-flowered or fasciculate, often grouped into panicles; bracts often inconspicuous, sometimes forming calyxlike involucre, or large and brightly colored
. Flowers bisexual
, rarely unisexual
or polygamous, actinomorphic
. Perianth constricted
beyond the ovary, base
persistent
, closely enclosing ovary which appears inferior, limb petaloid
beyond constriction, tubular
, funnelform
, or campanulate
, apex 5-10-lobed, lobes
plicate
or valvate
in bud, persistent or caducous
. Disk absent. Stamens (1-) 3-5(-many), hypogynous, free
or connate
at base, involute
in bud; anthers
2-loculed, dehiscence longitudinal
. Ovary superior, 1-loculed; ovule 1. Style 1; stigma globose
. Fruit an achenelike anthocarp enclosed by persistent perianth, ribbed
or winged
, often glandular
. Seed 1; endosperm present; embryo straight or curved
.
About 30 genera and 300 species: tropics and subtropics, mainly in tropical
America; six genera (two introduced
) and 13 species (one endemic, three introduced) in China.[1]
Genus Abronia
Herbs, annual
or perennial
, sometimes cespitose or appearing acaulescent
, usually viscid-pubescent, from slender to stout taproot
(extensively rhizomatous
in Abronia bolackii). Stems prostrate
to erect
, unarmed
, without glutinous
bands
on internodes. Leaves usually basal and cauline (all basal in A. bigelovii and A. nana), unequal in each pair, petiolate
; blade
± thick and succulent, base
usually asymmetric
. Inflorescences axillary
(appearing scapose
in A. bigelovii and A. nana), pedunculate
, capitate clusters
, with peripheral flowers usually opening first; receptacle slightly rounded
to conic, without pedicel-like projections; bracts persistent
, not accrescent
, 5-10, distinct
, forming involucre, lanceolate to broadly ovate
, thinly papery
or scarious
, translucent
, occasionally thin and green. Flowers bisexual
, chasmogamous
; perianth radially symmetric
, funnelform
or salverform
, constricted
distal to ovary, abruptly expanded to 5-lobed limb; stamens 5-9, included
; styles included; stigmas linear
. Fruits winged
or not, usually fusiform
or turbinate
, in profile
rhombic
, cordate, or obdeltate, coriaceous
, glabrate
to viscid
puberulent
; wings 2-5, opaque
, subtly veined, not or only slightly extending beyond apex or base of body, distal margins sometimes dilated
and flattened, broader than lamina, interior hollow, forming cavity
, or filled with spongy
tissue
; sulci smooth
or slightly rugose
.
Species ca.
20: North America, Mexico.
Mature
to near-mature fruits are usually required for identification of Abronia species because of the variation
of vegetative
structures within each taxon
. Abronia appears to be in a state of active
evolution. Cross-pollination
readily occurs in the greenhouse, producing a variety of hybrids. Hybridization occasionally occurs in the field
.[2]
Physical Description
Species Abronia pogonantha
Plants annual . Stems decumbent to ascending , little to moderately branched, elongate , glandular-pubescent . Leaves: petiole 0.8-4 cm; blade ovate to oblong-ovate, 2-5 × 1.5-3.5 cm, margins entire to ± sinuate and undulate , surfaces glandular-pubescent. Inflorescences: peduncle longer than subtending petiole; bracts lanceolate to broadly ovate, 4-9 × 1.8-6 mm, papery , puberulent to glandular-pubescent, often villous basally; flowers 12-24. Perianth: tube rose red, at least basally, 10-20 mm, limb white to pink, 6-8 mm diam. Fruits winged , broadly cordate or ± round in profile , 4-6 × 4-6 mm, coriaceous , apex deeply notched with small beak ; wings 2(-3), infrequently partially folded together, without dilations, interior spongy . [source]
Habit: Forb/herb
Flowers: Bloom Period: February, March, April, May, June, July. • Flower Color: lavender, pink, violet
Size/Age/Growth
Size: 6-12" tall.
Habitat
Sandy soils, Joshua tree
desert scrub
; 100-1600 m
[3].
Biology
Reproduction
Duration: Annual
Growth
Culture: Space 15-18" apart.
Soil: Minimum pH: 5.6 • Maximum pH: 6.5
Sunlight: Sun Exposure: Full Sun .
Moisture: Drought Tolerance: High
Temperature: Cold Hardiness: 8a, 8b, 9a, 9b, 10a, 10b. (map)
Taxonomy
- Domain:
Eukaryota
(
)
- Whittaker & Margulis,1978
- eukaryotes
- Kingdom:
Plantae
(
)
- Haeckel, 1866
- Plants
- Subkingdom:
Viridaeplantae
(
)
- Cavalier-Smith, 1981
- Phylum:
Tracheophyta
(
)
- Sinnott, 1935 Ex Cavalier-Smith, 1998
- Vascular Plants
- Subphylum:
Euphyllophytina
(
)
- Infraphylum:
Radiatopses
(
)
- Kenrick & Crane, 1997
- Class:
Magnoliopsida
(
)
- Brongniart, 1843
- Dicotyledons
- Subclass:
Caryophyllidae
(
)
- Takhtajan, 1967
- Superorder:
Caryophyllanae
(
)
- Takhtajan, 1967
- Order:
Caryophyllales
(
)
- Perleb, 1826
- Suborder:
Phytolaccineae
(
)
- Family:
Nyctaginaceae
(
)
- A.l. De Jussieu, 1789, Nom. Cons.
- Four O'clock Family
- Family:
Nyctaginaceae
(
- Suborder:
Phytolaccineae
(
- Order:
Caryophyllales
(
- Superorder:
Caryophyllanae
(
- Subclass:
Caryophyllidae
(
- Class:
Magnoliopsida
(
- Infraphylum:
Radiatopses
(
- Subphylum:
Euphyllophytina
(
- Phylum:
Tracheophyta
(
- Subkingdom:
Viridaeplantae
(
- Kingdom:
Plantae
(
Notes
Name
Status: Accepted Name
.
Publishing author
: Heimerl Publication
: Bot. Jahrb. Syst. 11: 87, pl. 2 1889An accepted name in the RHS Horticultural Database.
Similar Species
Members of the genus Abronia
ZipcodeZoo has pages for 120 species, subspecies, varieties, forms, and cultivars in this genus. Here are just 100 of them:
A. acutalata · A. alba (Whiteflower Sand Verbena) · A. alba platyphylla · A. alba var. platyphylla · A. alba variabilis · A. alpina (Ramshaw Meadows Sand Verbena) · A. ameliae (Amelia´s Sand-Verbena) · A. ammophila (Wyoming Sand Verbena) · A. angulata · A. angustifolia (Narrow-Leaf Sandverbena) · A. arenaria · A. argillosa (Clay Sand Verbena) · A. arizonica · A. bakeri · A. bigelovii (Galisteo Sand Verbena) · A. bigelovii Heimerl 'Galisteo Sand Verbena' · A. bolackii (Bolacks Sand Verbena) · A. breviflora · A. californica · A. carletoni (Carleton's Sand Verbena) · A. carletonii (Carleton's Sand Verbena) · A. carnea · A. carnea var. carnea · A. carnea var. wootonii · A. carterae · A. cheradophila · A. covillei · A. crux-maltae · A. cycloptera · A. deppei · A. elliptica (Fragrant White Sand Verbena) · A. exalata · A. fallax · A. fendleri · A. fragrans (Fragrant White Sand-Verbena) · A. fragrans elliptica · A. fragrans glaucescens · A. fragrans var. pterocarpa · A. glabra · A. glabrifolia · A. glauca · A. glaucescens · A. gracilis (Slender Sand Verbena) · A. gracilis subsp. platyphylla · A. insularis (Island Sand Verbena) · A. lanceolata · A. latifolia (Coastal Sand Verbena) · A. latiuscula · A. lobatifolia · A. lythrochiladel · A. macrocarpa (Large-Fruited Sand Verbena) · A. maritima (Red Sand Verbena) · A. maritima maritima · A. mellifera (Honey-Scent Sand Verbena) · A. menziesii · A. micrantha · A. micrantha pedunculata · A. minor (Little Sand Verbena) · A. nana (Dwarf Sand Verbena) · A. nana covillei (Coville's Dwarf Abronia) · A. nana S.Watson var. harrisii Welsh (Harris' Dwarf Sand Verbena) · A. nana lanciformis · A. nana S.Watson var. nana S.Watson (Dwarf Sand Verbena) · A. nana var. covillei (Coville's Dwarf Sand Verbena) · A. nana var. harrisii (Harris' Dwarf Sand Verbena) · A. nana var. nana (Dwarf Sand Verbena) · A. nealleyi · A. nelsonii · A. neurophylla (Beach Sand Verbena) · A. nudata · A. ochoterenae · A. orbiculata · A. parviflora · A. pedunculata · A. pinetorum · A. pinetorum var. aurita · A. platyphylla (Broadleaf Sand Verbena) · A. pogonantha (Mojave Sand Verbena) · A. pumila · A. ramosa · A. robusta · A. rosea · A. rotundifolia · A. sparsifolia · A. speciosa · A. suksdorfii · A. taeniata graminea · A. taeniata taeniata · A. taeniatus · A. texana · A. torreyi · A. turbinata (Transmontane Sand Verbena) · A. turbinata var. marginata · A. umbellata (Pink Sand Verbena) · A. umbellata acutalata · A. umbellata alba · A. umbellata breviflora (Pink Sand Verbena) · A. umbellata 'Grandiflora' · A. umbellata 'Pink Sand Verbena' · A. umbellata platyphylla
More Info
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- Search using Specialized Databases: GenBank | Medline | Scirus | CISTI/CAL | Agricola Periodicals | Agricola Books
Further Reading
- A flora of California, by Willis Linn Jepson. San Francisco, Calif., Cunningham, Curtis & Welch, 1909- ENG url p. 453, p. 455.
- An illustrated flora of the Pacific States: Washington, Oregon, and California. Stanford University, Stanford University Press, 1923-[60] ENG url p. 108, p. 111, p. 112.
- Botanisches Zentralblatt; referierendes Organ für das Gesamtgebiet der Botanik. Jena [etc.]G. Fischer [etc.] GER url p. 124.
- Lu Dequan. 1996. Nyctaginaceae. In: Tang Changlin, ed., Fl. Reipubl. Popularis Sin. 26: 114.
- Galloway, L. A. 1975. Systematics of North American desert species of Abronia and Tripterocalyx. Brittonia 27: 328-347.
- Tillett, S. S. 1967. The maritime species of Abronia (Nyctaginaceae). Brittonia 19: 299-327.
Notes
Contributors
- "Abronia pogonantha". in Flora of North America Vol. 4 Page 61, 64, 66. Published by Oxford University Press. Online at EFloras.org.
- Bisby FA, Roskov YR, Orrell TM, Nicolson D, Paglinawan LE, Bailly N, Kirk PM, Bourgoin T, van Hertum J, eds (2008). Species 2000 & ITIS Catalogue of Life: 2008 Annual Checklist Taxonomic Classification. CD-ROM; Species 2000: Reading, U.K.
- Global Biodiversity Information Facility. Accessed November 12, 2007. http://www.gbif.org Mediated distribution data from 2 providers.
Data Sources
Accessed through GBIF Data Portal November 12, 2007:
- Berkeley Natural History Museums, University and Jepson Herbaria DiGIR provider
- USDA PLANTS, USDA PLANTS Database
Identifiers
- Biodiversity Heritage Library NamebankID: 2646075
- Catalogue of Life Accepted Name Code: ITS-19570
- Global Biodiversity Information Facility Taxonkey: 13730067
- Globally Unique Identifier: urn:lsid:ipni.org:names:604009-1
- Integrated Taxonomic Information System (ITIS) Taxonomic Serial Number (TSN): 19570
- International Plant Names Index (IPNI) ID: 101-2
- Natural Heritage Network Species Identifier: PDNYC010L0
- U.S.D.A. Plant Symbol: ABPO
- Zipcode Zoo Species Identifier: 71133
Footnotes
- Dequan Lu & Michael G. Gilbert "Nyctaginaceae". in Flora of China Vol. 5 Page 430. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org. [back]
- Leo A. Galloway "Abronia". in Flora of North America Vol. 4 Page 16, 61. Oxford University Press. Online at EFloras.org. [back]
- "Abronia pogonantha". in Flora of North America Vol. 4 Page 61, 64, 66. Oxford University Press. Online at EFloras.org. [back]
