Overview
|
Critically Endangered |
|
Common Names
Common Names in English:
Ko´oloa´ula, Ko'oloa 'ula, Ko'oloa'ula, Kooloaula, Ko`oloa`ula, Red Llima
Description
Family Malvaceae
Herbs, shrubs
, or less often trees
; indumentum usually with peltate scales
or stellate
hairs
. Leaves alternate, stipulate
, petiolate
; leaf blade
usually palmately veined, entire or various lobed
. Flowers solitary, less often in small cymes or clusters
, axillary
or subterminal
, often aggregated into terminal
racemes
or panicles, usually conspicuous
, actinomorphic
, usually bisexual
(unisexual
in Kydia) . Epicalyx
often present, forming an involucre around calyx, 3- to many lobed. Sepals 5, valvate
, free
or connate
. Petals 5, free, contorted, or imbricate, basally adnate
to base
of filament
tube
. Stamens usually very many, filaments connate into tube; anthers
1-celled. Pollen spiny
. Ovary superior, with 2-25 carpels, often separating from one another and from axis; ovules 1 to many per locule; style as many or 2 × as many as pistils, apex branched or capitate. Fruit a loculicidal capsule or a schizocarp, separating into individual mericarps, rarely berrylike when mature
(Malvaviscus) ; carpels sometimes with an endoglossum (a crosswise projection from back wall of carpel to make it almost completely septate
. Seeds often reniform
, glabrous
or hairy
, sometimes conspicuously so.
About 100 genera and ca.
1000 species: tropical
and temperate regions
of N and S Hemisphere; 19 genera (four introduced
) and 81 species (24 endemic, 16 introduced) in China.
Molecular studies have shown that the members
of the Bombacaceae, Malvaceae, Sterculiaceae, and Tiliaceae form a very well-defined monophyletic group that is divided
into ten also rather well-defined clades, only two of which correspond to the traditional families Bombacaceae and Malvaceae. Some of the remaining groups are included
entirely within either of the remaining families but others cut
across the traditional divide between the Sterculiaceae and Tiliaceae. A majority of authors
, most notably Bayer and Kubitzki (Fam. Gen. Vasc. Pl. 5: 225-311. 2003), has favored including everything within a greatly enlarged Malvaceae, and treating the individual clades as subfamilies. The alternative view
is that the individual clades should be treated as a series of ten families: Bombacaceae (Bombacoideae), Brownlowiaceae (Brownlowioideae), Byttneriaceae (Byttnerioideae), Durionaceae (Durionoideae), Helicteraceae (Helicteroideae), Malvaceae (Malvoideae), Pentapetaceae (Dombeyoideae), Sparrmanniaceae (Grewioideae), Sterculiaceae (Sterculioideae), and Tiliaceae (Tilioideae) (Cheek in Heywood et al.
, Fl.
Pl. Fam. World. 201-202. 2007) . For the present treatment, we prefer to retain the familiar, traditional four families, so as to maintain continuity
with the treatments in FRPS, and to await a consensus on the two alternative strategies for dealing with the very widely accepted clades.
The traditional Malvaceae coincides exactly with one of the major clades. The only possible problem is the relationship
with the Bombacaceae, which also has primarily 1-loculed anthers, and some authorities have suggested that the Bombacaceae should be included within the Malvaceae.
Members of the Malvaceae are important as fiber crops
(particularly cotton, Gossypium) . Young leaves of many species can be used as vegetables, and species of Abelmoschus and Hibiscus are grown as minor food crops. Many species have attractive flowers and an ever-increasing selection is grown as ornamentals
. Several have been cultivated for a very long time, particularly species of Hibiscus, and some of these are not known in the wild.[1]
Genus Abutilon
Herbs, subshrubs
, shrubs
, or small trees
. Stipules usually caducous
; leaf blade
usually entire (lobed
in A. pictum), palmately veined, base
cordate, margin
crenate
or serrate. Flowers axillary
or subterminal
, solitary, paired
or in small cymes, often aggregated into terminal
panicles. Epicalyx
absent. Calyx campanulate
, lobes
5. Corolla mostly yellow or orange (red in A. roseum), often with dark center, campanulate to wheel-shaped, rarely ± tubular
(A. pictum) ; petals 5, basally connate
and adnate
to filament
tube
. Anthers
many, clustered at filament tube apex. Ovary (5-) 7-20-loculed; ovules 2-9 per carpel; style branches as many as carpels. Fruit a schizocarp, often blackish when mature
, subglobose to hemispherical; mericarps (5-) 7-20, eventually dehiscent
, apex rounded
or acute, sometimes 2-awned, pericarp leathery. Seeds reniform
, glabrous
or slightly pubescent
.
About 200 species: mostly in tropics and subtropics; nine species (three endemic, one introduced
) in China.
Several species have become widespread tropical
weeds
. One species, Abutilon pictum, is widely grown as an ornamental
.[2]
Physical Description
Habit: Shrub
Flowers: Bloom Period: January, February, March, April, May, July, August, September, October, November, December. • Flower Color: magenta, maroon, mauve , pale pink, pink, rose
Size/Age/Growth
Size: 6-8' tall.
Habitat
Typically found at an altitude of 0 to -4,496 meters (0 to -14,751 feet).[3]
Ecology: A perennial
shrub
of dry shrublands, including ones that are now dominated by alien
plant species. Gulches, plateau
lands, and old lava flows
.[4]
List of Habitats
:1.5Forest - Subtropical/Tropical Dry
3.5Shrubland - Subtropical/Tropical Dry
Biology
Reproduction
Duration: Perennial
Growth
Culture: Space 6-8' apart.
Sunlight: Sun Exposure: Full sun .
Taxonomy
- Domain:
Eukaryota
(
)
- Whittaker & Margulis,1978
- eukaryotes
- Kingdom:
Plantae
(
)
- Haeckel, 1866
- Plants
- Subkingdom:
Viridaeplantae
(
)
- Cavalier-Smith, 1981
- Phylum:
Tracheophyta
(
)
- Sinnott, 1935 Ex Cavalier-Smith, 1998
- Vascular Plants
- Subphylum:
Euphyllophytina
(
)
- Infraphylum:
Radiatopses
(
)
- Kenrick & Crane, 1997
- Class:
Magnoliopsida
(
)
- Brongniart, 1843
- Dicotyledons
- Subclass:
Dilleniidae
(
)
- Takhtajan, 1967
- Superorder:
Malvanae
(
)
- Takhtajan, 1967
- Order:
Malvales
(
)
- Dumortier, 1829
- Family:
Malvaceae
(
)
- Adans., 1763, Nom. Cons.
- Mallow Family
- Subfamily:
Malvoideae
(
)
- Tribe:
Abutilieae
(
)
- Genus:
Abutilon
(
)
- P. Miller, 1754
- Indian Mallow
- Specific epithet:
menziesii
- Seem.
- Botanical name: - Abutilon menziesii
- Specific epithet:
menziesii
- Seem.
- Genus:
Abutilon
(
- Tribe:
Abutilieae
(
- Subfamily:
Malvoideae
(
- Family:
Malvaceae
(
- Order:
Malvales
(
- Superorder:
Malvanae
(
- Subclass:
Dilleniidae
(
- Class:
Magnoliopsida
(
- Infraphylum:
Radiatopses
(
- Subphylum:
Euphyllophytina
(
- Phylum:
Tracheophyta
(
- Subkingdom:
Viridaeplantae
(
- Kingdom:
Plantae
(
Notes
Name
Status: Accepted Name
. Latest taxonomic
scrutiny: 15-Mar-2000
Place of publication
: Fl.
vit. 15. 1865
Name verified on 19-Apr-1994 by ARS Systematic Botanists. Last updated: 03-Jan-2002
Similar Species
Members of the genus Abutilon
ZipcodeZoo has pages for 692 species, subspecies, varieties, forms, and cultivars in this genus. Here are just 100 of them:
A. - · A. abutiloides (American Indian-Mallow) · A. abutilon · A. acerifolium · A. affine · A. agnesae · A. albescens · A. albidum · A. album · A. alii · A. 'Alpha Glory' · A. ambiguum · A. americanum · A. 'Amiti' · A. amoenum · A. amplexifolium · A. amplissimum · A. amplum · A. 'Amsterdam' · A. anderssonianum · A. andrewsianum · A. andrieuxii · A. anglosomaliae · A. angulatum · A. angulatum var. macrophyllum · A. anodoides · A. 'Apollo' (Chinese Bell Flower) · A. appendiculatum · A. 'Apricot' (Flowering Maple) · A. 'Apricot Belle' · A. 'Apricot Glow' · A. arborescens · A. arboreum · A. arenarium · A. arequipense · A. aristulosum · A. arnottianum · A. 'Ashford Red' (Flowering Maple) · A. asiaticum (Asian Indian Mallow) · A. asiaticum var. albescens · A. asiaticum var. australiense · A. asiaticum var. subasperum · A. asiaticum var. supraviride · A. asperifolium · A. atropurpureum · A. attenuatum · A. aurantiacum · A. aureum · A. auritum (Asian Indian Mallow) · A. australe · A. australe var. malvifolium · A. austroafricanum · A. avicennae · A. avicennae var. chinense · A. 'Baby Orange' (Flowering Maple) · A. badium · A. bakeranum · A. bakeri · A. bakerianum · A. balansae · A. 'Ballerina' · A. barrancae · A. 'Bartley Schwartz' (Chinese Bell Flower) · A. bastardioides · A. bathiei · A. bedfordianum · A. bedfordianum discolor · A. bedfordianum subsp. concolor · A. bedfordianum subsp. discolor · A. behrianum · A. 'Bella Coral' (Flowering Maple) · A. 'Bella Pink' (Flowering Maple) · A. 'Bella Salmon Shades' · A. 'Bella Select Mix' (Flowering Maple) · A. Bella Series · A. 'Bella Vanilla' (Flowering Maple) · A. benadirense · A. 'Benary's Giant' · A. benedictum · A. benense · A. berlandieri (Berlandier's Indian Mallow) · A. 'Bessie Schulson' · A. betschuanicum · A. bicolor · A. bidentatum · A. bidentatum var. forrestii · A. blandum · A. blepharocarpum · A. 'Bloomsbury Can-can' · A. 'Bloomsbury Rose' · A. 'Boule de Neige' · A. bracteosum · A. braunii · A. brenesii · A. bridgesii · A. buchii · A. burandtii · A. bussei · A. cabrae · A. caldemni
More Info
- Search for Pictures: images.google.com
- Search for Scholarly Articles: Google Scholar
- Search using Scientific Name and Vernacular Names: All the Web | AltaVista Canada | AltaVista | Excite | Google | HotBot | Lycos
- Search using Specialized Databases: GenBank | Medline | Scirus | CISTI/CAL | Agricola Periodicals | Agricola Books
Further Reading
- (2002). Hawaiian Native Plant Genera. ¬ Gerald D. Carr, University of Hawaii Botany Department. http://www.botany.hawaii.edu/faculty/carr/natives.htm. Accessed: 2002.
- Bornhorst, H.L.; Rauch, F.D. 1994. Native Hawaiian Plants for Landscaping, Conservation, and Reforestation. Honolulu, HI: University of Hawaii--Hawaii Institute of Tropical Agriculture and Human Resources. 18p.
- Ellshoff, Z.E. 1991. The Rarest Hawaiian Members of the Hibiscus Family. National Tropical Botanical Garden: The Bulletin. 21, 3: 7-12.
- Gustafson, Robert J. Hawaii's Unique and Vanishing Flora: A Photographic Exhibition. [Web site] ¬ The Natural History Museum of Los Angeles County Foundation. http://www.nhm.org/research/botany/Hawaii_Vanishing_Flora/home.html. Accessed: 2002.
- Kartesz, J.T. 1999. A synonymized checklist of the vascular flora of the U.S., Canada, and Greenland. In: Kartesz, J.T.; Meacham, C.A., editors. Synthesis of the North American Flora, Version 1.0. North Carolina Botanical Garden. Chapel Hill, NC.
- NatureServe. 2003. NatureServe Explorer: An online encyclopedia of life [web application]. Version 1.8. NatureServe, Arlington, Virginia. (Accessed: October 17, 2003).
- U.S. Fish and Wildlife Service. 2003. Rare plant database. Unpublished.
- USFWS. 1976. Proposed Endangered Status for 1700 U.S. Plants. Federal Register. 41: 24523-24572.
- USFWS. 1985. Public Hearings and Reopening of Comment Period on Proposed Endangered Status for Abutilon menziesii (ko`oloa 'ula), Hibiscadelphus distans (Kauai hau kuahiwi), Mezoneuron kavaiense (uhiuhi), and Scaevola coriacea (dwarf naupaka). Federal Register. 50, 202: 42196-42197.
- USFWS. 1995. Lana`i Plant Cluster Recovery Plan. Portland, Oregon: U.S. Fish and Wildlife Service. p.138.
- United States Fish and Wildlife Service. U.S. Fish & Wildlife Service. Species Information: threatened and endangered animals and plants - on-line resource. (US Fish & Wildl List)
- Wagner, W. L. et al. 1990. Manual of the flowering plants of Hawai'i. (F Hawaii)
- Wagner, W. L. et al. 1999. Hawaiian vascular plants at risk:1999. Bishop Mus. Occas. Pap. 60:40.
- Wagner, W., Herbst, D. and Sohmer S. 1999. Manual of the Flowering Plants of Hawaii. Bernice Pauahi Bishop Museum Special Publication 91: 1-1918.
- Wagner, W.L.; Herbst, D.R. 2002. Supplement to the Manual of the Flowering Plants of Hawai'i. University of Hawaii Press.
- Yoshinaga, Alan. (2002). Seed storage characteristics of Hawaiian species. [Web site] University of Hawaii, Honolulu, Hawaii 96848. http://www.hawaii.edu/scb/seed/seedtabl.html. Accessed: 2002.
- Feng Kuo-mei. 1984. Malvaceae. In: Feng Kuo-mei, ed., Fl. Reipubl. Popularis Sin. 49(2): 1-102.
- Feng Kuo-mei. 1984. Malvaceae. In: Feng Kuo-mei, ed., Fl. Reipubl. Popularis Sin. 49(2): 1-102.
Notes
Contributors
- Bruegmann, M.M. & Caraway, V. 2003. In IUCN 2008. 2008 IUCN Red List of Threatened Species. IUCNRedList.org. Downloaded July 19, 2008.
- Global Biodiversity Information Facility. Accessed February 02, 2008. http://www.gbif.org Mediated distribution data from 4 providers.
- USDA, ARS, National Genetic Resources Program. Germplasm Resources Information Network - (GRIN) [Online Database]. National Germplasm Resources Laboratory, Beltsville, Maryland. URL (April 24, 2008)
Data Sources
Accessed through GBIF Data Portal February 02, 2008:
- Bernice Pauahi Bishop Museum, Bishop Museum Natural History Specimen Data
- USDA PLANTS, USDA PLANTS Database
Identifiers
- Biodiversity Heritage Library NamebankID: 3465155
- Catalogue of Life Accepted Name Code: ITS-21680
- GRIN Nomen Number: 320088
- Integrated Taxonomic Information System (ITIS) Taxonomic Serial Number (TSN): 21680
- International Plant Names Index (IPNI) ID: 558365-1
- IUCN ID: 44060
- Natural Heritage Network Species Identifier: PDMAL020C0
- U.S. Fish and Wildlife Service Species Identifier: Q1SK
- U.S.D.A. Plant Symbol: ABME2
- Zipcode Zoo Species Identifier: 1856
Footnotes
- Ya Tang, Michael G. Gilbert & Laurence J. Dorr "Malvaceae". in Flora of China Vol. 12 Page 240, 264,299, 302. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org. [back]
- "Abutilon". in Flora of China Vol. 12 Page 265, 275. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org. [back]
- Mean = -1,301.530 meters (-4,270.112 feet), Standard Deviation = 1,311.530 based on 17 observations. Altitude information for each observation from British Oceanographic Data Centre. [back]
- Bruegmann, M.M. & Caraway, V. 2003. In IUCN 2008. 2008 IUCN Red List of Threatened Species. IUCNRedList.org. Downloaded July 19, 2008. [back]
