Achyranthes aspera

• Common Names
• Description
• Biology
• Taxonomy
• Similar Species
• More Info
• Further Reading
• Notes

Common Names

Click on the language to view common names.

Description

Family Amaranthaceae

Herbs, clambering subshrubs , shrubs , or lianas. Leaves alternate or opposite, entire, exstipulate . Flowers small, bisexual or unisexual , or sterile and reduced, subtended by 1 membranous bract and 2 bracteoles, solitary or aggregated in cymes. Inflorescences elongated or condensed spikes (heads ), racemes , or thyrsoid structures of varying complexity. Bracteoles membranous or scarious . Tepals 3-5, membranous, scarious or subleathery, 1-, 3-, 5-, or 7(-23) -veined. Stamens as many as tepals and opposite these, rarely fewer than tepals; filaments free , united into a cup at base or ± entirely into a tube , filament lobes present or absent, pseudostaminodes present or absent; anthers (1- or) 2-loculed, dorsifixed , introrsely dehiscent . Ovary superior, 1-loculed; ovules 1 to many; style persistent , short and indistinct or long and slender; stigma capitate, penicillate , 2-lobed or forming 2 filiform branches. Fruit a dry utricle or a fleshy capsule, indehiscent, irregularly bursting, or circumscissile. Seeds lenticular , reniform , subglobose, or shortly cylindric , smooth or verruculose .

About 70 genera and 900 species: worldwide; 15 genera (one introduced ) and 44 species (three endemic, 14 introduced) in China.

Morphology of the androecium, perianth (tepals), and the inflorescence has traditionally been used to circumscribe genera and tribes . Pseudostaminodia are interstaminal appendages with variously shaped apices. Filament appendages are the lateral appendages of filaments (one on each side) . The basic structure of the inflorescence is the cyme (branchlets arising from the bracteole axils, the bracteoles serving as bracts for upper flowers), which can be reduced to one flower with two bracteoles and a bract. Units of dispersal vary considerably (capsules opening with lower part persistent, flower and bracteoles falling together, or cymose partial inflorescences breaking off above bract) and can be characteristic for genera. Several genera possess long trichomes serving dispersal at the base of the tepals.[1]

Genus Achyranthes

Herbs, perennial or annual . Stems erect to ascending . Leaves opposite, petiolate ; blade elliptic , ovate to orbiculate, or broadly rhombate, margins entire. Inflorescences terminal and axillary , pedunculate , elongate , many-flowered, simple spikes or few-branched panicles; flowers crowded together at tips , becoming more widely spaced toward base . Flowers bisexual , often becoming deflexed with age; tepals 4 or 5, basally connate , without ornamentation, coriaceous , becoming indurate in fruit, ± glabrous ; filaments basally connate into short tubes or cups ; anthers 4-locular; pseudostaminodes 5; ovary obovoid or turbinate ; ovule 1; style elongate; stigma 1, capitate. Utricles enclosed by and falling with indurate tepals, elliptic or cylindric , membranous, indehiscent. Seeds 1, inverted , obovoid or ovoid , smooth .

Species 8-12: c and se United States, Mexico, West Indies, Central America, South America, tropical , subtropical , and warm-temperate regions of the Old World.

The groups of plants referred to as Achyranthes and Alternanthera have been subject to considerable nomenclatural confusion, primarily because P. C. Standley (1915) designated Achyranthes repens Linnaeus as the lectotype species of Achyranthes. As a result, species that had been placed in Achyranthes were transferred to Centrostachys Wallich, and species that had been in Alternanthera were transferred to Achyranthes. A. A. Bullock (1957; see also R. Melville 1958) showed that Standley's lectotypification was incorrect and that the type species of Achyranthes is Achyranthes aspera Linnaeus. The generic concepts of Achyranthes and Alternanthera then returned to those prior to 1915.[2]

Physical Description

Habitat

Biology

Reproduction

Duration: Annual , Perennial

Taxonomy

Unambiguous Synonyms

1. Achyranthes argentea Lam.
2. Achyranthes aspera obtusifolia (Lam.) Griseb.
3. Achyranthes obtusifolia Lamarck
4. Centrostachys aspera (L.) Standl.

Notes

Publishing author : Standl. Publication : J. Wash . Acad. Sci. 1915, v. 75, in obs. Publishing author: Standl. Publication: J. Wash. Acad. Sci. 5: 75 1915 Basionym author: (L.) Publishing author: Lam. Publication: Encyc. i. 545 Publishing author: Mill . Publication: Gard. Dict., ed. 8. [unpaged] Achyranthes no. 2 1768 Basionym author: (L.) Name Status: Accepted Name . Latest taxonomic scrutiny: 15-Mar-2000

Place of publication: Sp. pl. 1:204. 1753, nom. cons.

Name verified on 12-Mar-2004 by ARS Systematic Botanists. Last updated: 29-Oct-2007

Similar Species

Members of the genus Achyranthes

ZipcodeZoo has pages for 216 species, subspecies, varieties, forms, and cultivars in this genus. Here are just 100 of them:

A. abyssinica · A. acuminata · A. alba · A. albida · A. alopecuroides · A. alternifolia · A. altissima · A. amabilis · A. amaranthoides (Jiang Guo Xian) · A. ancistrophora · A. angustifolia · A. annua · A. aphylla · A. aquatica · A. arborescens · A. argentea · A. aspera (Devil´s Horsewhip) · A. aspera obtusifolia · A. aspera var. argentea · A. aspera var. aspera (Devil's Horsewhip) · A. aspera var. canescens · A. aspera var. indica · A. aspera var. obtusifolia · A. aspera var. porphyristachya · A. aspera var. pubescens (Devil's Horsewhip) · A. aspera var. rubro-fusca · A. aspera var. sicula · A. aspera var. simplex · A. aspera var. velutina · A. atollensis (Hawai'i Chaff Flower) · A. atra · A. atropurpurea · A. australis · A. avicularis · A. axillaris · A. baccata · A. bangii · A. bengalensis · A. benthamii · A. bettzickiana · A. bidentata (Pig's Knee) · A. bidentata f. rubra · A. bidentata var. japonica · A. bidentata var. villosa · A. biemilleri · A. boliviana · A. borbonica · A. brachiata · A. brasiliana · A. breviflora · A. calceolata · A. calea · A. canescens · A. capitata · A. capituliflora · A. carnosa · A. carsonii · A. chacoensis · A. chinensis · A. ciliata · A. conferta · A. cordata · A. cordobensis · A. corymbosa · A. costaricensis · A. coynei · A. crassifolia · A. crispa · A. cyclophylla · A. cylindrica · A. debilis · A. decumbens · A. diandra · A. dichotoma · A. digera · A. dioica · A. echinata · A. echinocephala · A. elegantissima · A. elongata · A. excelsa · A. fasciculata · A. fauriei · A. ferruginea · A. ficoidea · A. filifolia · A. fischeri · A. flavicoma · A. frumentacea · A. frutescens · A. fruticosa · A. fruticosa var. pubescens · A. galapagensis · A. geminata · A. geniculata · A. glauca · A. glaucescens · A. globosa · A. glomerata · A. gracilis

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Further Reading

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Notes

Contributors

• "Achyranthes aspera". in Flora of North America Vol. 4 Page 435, 436. Published by Oxford University Press. Online at EFloras.org.
• Bisby, F.A., Y.R. Roskov, M.A. Ruggiero, T.M. Orrell, L.E. Paglinawan, P.W. Brewer, N. Bailly, J. van Hertum, eds (2007). Species 2000 & ITIS Catalogue of Life: 2007 Annual Checklist. Species 2000: Reading, U.K.
• Brands, S.J. (comp.) 1989-2006. Systema Naturae 2000. The Taxonomicon. Universal Taxonomic Services, Amsterdam, The Netherlands. Accessed April 21, 2007.
• Global Biodiversity Information Facility. Accessed December 13, 2007. http://www.gbif.org Mediated distribution data from provider.
• Kirk P.M., 14-Sep-2005 (from Species Fungorum, Sep 2006).
• The International Plant Names Index. Accessed Jan 19, 2007.
• USDA, ARS, National Genetic Resources Program. Germplasm Resources Information Network - (GRIN) [Online Database]. National Germplasm Resources Laboratory, Beltsville, Maryland. URL (April 25, 2008)

Data Sources

Accessed through GBIF Data Portal November 15, 2007:

• Australian National Herbarium
• , Australian National Herbarium
• GBIF-Spain, Institut Botanic de Barcelona, BC
• GBIF-Spain, Jardín Botánico de Córdoba: Herbarium COA
• Harvard University Herbaria, Harvard University Herbaria
• Herbarium of the University of Aarhus, The AAU Herbarium Database
• Herbier de la Guyane, Herbier de la Guyane
• Instituto Nacional de Biodiversidad
• , Biodiversidad de Costa Rica
• Israel Nature and Parks Authority, Israel Nature and Parks Authority
• Missouri Botanical Garden, Missouri Botanical Garden
• National Herbarium of New South Wales, NSW herbarium collection
• National Herbarium of New South Wales, Plants of Papua New Guinea
• Royal Botanic Gardens, Kew, Royal Botanic Gardens, Kew
• Taiwan Biodiversity Information Facility, Magnoliophyta
• USDA PLANTS, USDA PLANTS Database
• University of Alabama Biodiversity and Systematics, Herbarium

Identifiers

• Biodiversity Heritage Library NamebankID: 2646922
• Catalogue of Life Accepted Name Code: ITS-20756
• Global Biodiversity Information Facility Taxonkey: 13740634
• Globally Unique Identifier: urn:lsid:ipni.org:names:85335-3
• GRIN Nomen Number: 401486
• Integrated Taxonomic Information System (ITIS) Taxonomic Serial Number (TSN): 20756
• International Plant Names Index (IPNI) ID: 58663-1
• Natural Heritage Network Species Identifier: PDAMA01010
• U.S.D.A. Plant Symbol: ACASP
• Zipcode Zoo Species Identifier: 18478

Footnotes

1. Bojian Bao, Thomas Borsch & Steven E. Clemants "Amaranthaceae". in Flora of China Vol. 5 Page 415. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org. [back]
2. Kenneth R. Robertson "Achyranthes". in Flora of North America Vol. 4 Page 406, 435. Oxford University Press. Online at EFloras.org. [back]

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