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Adiantum capillus-veneris

(Southern maidenhair Fern)

Common Names

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Click on the language to view common names.

Common Names in Danish:

ægte Venushår, Venushår

Common Names in Dutch:

Europees Venushaar

Common Names in English:

Common Maidenhair, Common Maidenhair Fern, Maidenhair Fern, Southern maidenhair Fern, Southern Maidenhair, Southern Maidenhair Fern, Venus Hair, Venus Hairfern, Venus Maidenhair, Venus Maidenhair Fern, Venus´ Hair Fern, Venus's-Hair Fern

Common Names in Finnish:

Venuksenhiussaniainen

Common Names in French:

Capillaire Cheveux-De-Vénus, Capillaire De Montpellier, Cheveux De Vénus

Common Names in German:

Frauenhaarfarn, Venushaarfarn

Common Names in Italian:

Capelvenere Comune

Common Names in Spanish:

Adianto, Capilera, Culantrillo De Pozo

Description

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Genus Adiantum

Plants terrestrial or on rock. Stems short- to long-creeping or suberect, branched; scales deep tawny yellow to dark reddish brown [black], concolored or bicolored , linear-lanceolate to lanceolate, margins entire, erose-ciliate, or minutely dentate . Leaves monomorphic to somewhat dimorphic , densely clustered to closely spaced [distant ], 15--110 cm. Petiole chestnut brown to dark purple or blackish, with single groove adaxially, glabrous , hispid , or strigose , with 1 or 2 vascular bundles . Blade lanceolate, ovate , trowel-shaped, or fan-shaped, 1--4(--9) -pinnate proximally, membranaceous to papery , both surfaces commonly glabrous (2 species with scattered hairs ), adaxially dull or shiny, not striate ; rachis straight or flexuous . Ultimate segments subsessile to short-stalked (stalks terminating in cupulelike swelling at base of pinna in A. tenerum ), round, fan-shaped, rhombic , or oblong , 3--29 mm wide; base truncate to cuneate, free from costa; stalk dark, often lustrous ; fertile segments with marginal lobes recurved to form false indusia. Veins of ultimate segments conspicuous , free, ± dichotomously forking near base and well above segment base [anastomosing in a few tropical species], parallel distally. False indusia light gray-green or brown to dark brown, narrow, 0.6--1 mm wide, marginal, concealing sporangia until sporangia dehisce. Sporangia submarginal, borne along or sometimes also between veins on abaxial surface of false indusium, paraphyses and glands absent. Spores yellow or yellowish brown, tetrahedral-globose, trilete, rugulate to rugose or tuberculate , equatorial ridge absent. x = 29, 30.

Species ca. 150--200: nearly worldwide except at latitudes greater than 60°.

Most diverse in Andean South America, Adiantum is primarily a tropical genus; of the nine species occurring in the flora , A. melanoleucum, A. tenerum, and A. tricholepis are strictly subtropical . Adiantum hispidulum occurs only as an escape from cultivation. The genus is absent from dry areas in the interior of the continent.

Adiantum is a very clearly circumscribed genus of ferns, the character state "sporangia borne on abaxial surface of false indusium" being both necessary and sufficient to define it. Within this large and widespread genus, however, species relationships are mostly unknown. An evolutionary classification of the group is indeed much needed (R. M. Tryon and A. F. Tryon 1982).[1]

Physical Description

Species Adiantum capillus-veneris

Stems short-creeping; scales golden brown to medium brown, concolored, iridescent , margins entire or occasionally with single broad tooth near base . Leaves lax-arching or pendent, closely spaced, 15--75 cm. Petiole 0.5--1.5 mm diam., glabrous , occasionally glaucous. Blade lanceolate, pinnate, 10--45 × 4--15 cm, glabrous, gradually reduced distally; proximal pinnae 3(--4) -pinnate; rachis straight to flexuous , glabrous, not glaucous. Segment stalks 0.5--3.5 mm, dark color extending into segment base. Ultimate segments various, generally cuneate or fan-shaped to irregularly rhombic (plants in American southwest occasionally with segments nearly round), about as long as broad; base broadly to narrowly cuneate; margins shallowly to deeply lobed , incisions 0.5--7 mm, occasionally ± laciniate , sharply denticulate in sterile segments; apex rounded to acute. Indusia transversely oblong or crescent-shaped, 1--3(--7) mm, glabrous. Spores mostly 40--50 µm diam. 2 n = 120. Sporulating spring--summer. [source]

No evident pattern to morphologic variation in the species is discernible, although a number of segregate species and infraspecific taxa have been recognized within North American Adiantum capillus-veneris. In the Eastern Hemisphere, the species is diploid, with 2 n = 60 (I. Manton 1950). Several tetraploid counts have been reported from North America (W. H. Wagner Jr. 1963). Spore-measurement data suggest, however, that the polyploid cytotype may not be widely distributed. Further investigation is needed to determine whether Adiantum capillus-veneris populations in North America are conspecific with those in Eurasia and Africa. [source]

Habit: Forb/herbGrowth Form: RhizomatousShape and Orientation: Erect

Flowers: Bloom Period: Summer • Flower Color: White • Flower Conspicuous: No

Seeds: Seed per Pound: 3000000 • Seed Spread Rate: Slow • Seedling Vigor: Medium • Fruit/Seed Abundance: Medium • Fruit/Seed Color: Brown • Fruit/Seed Conspicuous: No • Cold Stratification Required: No

Foliage: Foliage Color: Green • Foliage Porosity Summer: Porous • Foliage Porosity Winter: Porous • Foliage Texture: Fine • Fall Conspicuous: No • Leaf Retention: No

Size/Age/Growth

Active Growth Period: Spring and Summer • Growth Rate: Moderate • After Harvest Regrowth Rate: Slow • Mature Height (feet): 1.7 • Size: 18-24" tall. • Vegetative Spread Rate: Slow • Lifespan: Lifespan

Habitat

Moist calcareous cliffs , banks, and ledges along streams and rivers , walls of lime sinks, canyon walls (in the American southwest), around foundations, on mortar of storm drains ; 0--2500 m [2].

Biology

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Reproduction

Duration: PerennialCoppice Potential: No • Progagated by Bulbs: No • Propagated by Bare Root: Yes • Propagated by Container: No • Propagated by Corms: No • Propagated by Cuttings: No • Propagated by Seed: Yes • Propagated by Sod: No • Propagated by Sprigs: No • Propagated by Tubers: No • Fruit/Seed Period Begin: Summer • Fruit/Seed Period End: FallFruit/Seed Persistence: No

Growth

Culture: Space 12-15" apart.

Soil: Adapted to Medium Textured: Adapted to Medium Textured Soils • Adapted to Coarse Textured Soils: No • Anaerobic Tolerance: Medium • Salinity Tolerance: None • CaCO3 Tolerance: High • Minimum pH: 6.0 • Maximum pH: 8.0 • Fertility Requirement: Low

Sunlight: Sun Exposure: Partial to Full Shade. • Shade Tolerance: Tolerant

Moisture: Drought Tolerance: None • Minimum Precipitation: 10 • Maximum Precipitation: 60 • Moisture Use: High

Temperature: Minimum Temperature (F): -33°F. • Minimum Frost Free Days: 120 • Cold Hardiness: 7a, 7b, 8a, 8b, 9a, 9b, 10a, 10b. (map)

Taxonomy

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Unambiguous Synonyms

  1. A. capillus-veneris var. protrusum Fernald
  2. A. capillus-veneris var. rimicola (Slosson) Fernald
  3. Adiantum capillus-veneris var. modestum (L. Underwood) Fernald
  4. Adiantum capillus-veneris var. protrusum Fern.
  5. Adiantum modestum Underwood


Notes

Publishing author : Fernald Publication : Rhodora 52: 203 1950 Publishing author: Und. Publication: Bull . Torr. Cl. 28. 46. 1901 1901 Name Status: Accepted Name . Latest taxonomic scrutiny: 15-Mar-2000

Place of publication: Sp. pl. 2:1096. 1753

Name verified on 14-Apr-1995 by ARS Systematic Botanists. Last updated: 18-Jul-2007

Similar Species

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Members of the genus Adiantum

ZipcodeZoo has pages for 641 species, subspecies, varieties, forms, and cultivars in this genus. Here are just 100 of them:

A. abscissum · A. achilleifolium · A. acinaciforme · A. acrocarpon · A. aculeatum · A. aculeolatum · A. acuminatum · A. adiantoides · A. aemulum · A. aethiopicum (Common Maidenhair Fern) · A. aethiopicum var. nodosa · A. aetiopi · A. affine · A. africanum · A. alarconianum · A. alatum · A. aleuticum (Aleutian Maidenhair-Fern) · A. aleuticum 'Dwarf Ecotype' · A. aleuticum 'Imbricatum' · A. aleuticum 'Japonicum' · A. aleuticum 'Laciniatum' · A. aleuticum 'Miss Sharples' · A. aleuticum 'Serpentine Ecotype' · A. aleuticum subpumilum · A. aleuticum 'Subpumilum' · A. amabile · A. amazonicum · A. amblyopteridium · A. amelianum · A. americanum · A. amoenum · A. amplum · A. anceps (Double Edge Maidenhair) · A. andicola · A. aneitense · A. angustatum · A. angustifolium · A. annamense · A. apalophyllum · A. arcanum · A. arcuatum · A. argutum · A. aristatum · A. asarifolium · A. asperum · A. assimile · A. atroviride · A. aubertii · A. auriculatum · A. australe · A. baenitzii · A. balansae · A. balfourii · A. bausei · A. bellum (Bermuda Maidenhair Fern) · A. bessoniae · A. birkenheadii · A. blumenavense · A. boliviense · A. bonatianum · A. bonii · A. bonplandii · A. borbonicum · A. boreale · A. brasiliense · A. braunii · A. breviserratum · A. caffrorum · A. cajennense · A. cajennense var. stenophyllum · A. calcareum · A. candatum · A. cantoniense · A. capense · A. capillaceum · A. capillaris-veneris · A. capillatum · A. capillis-veneris · A. capillium-veneris · A. capillius-veneris · A. capillus (Common Maidenhair) · A. capillus-junonis · A. capillus-venaris · A. capillus-venensis · A. capillus-veneri · A. capillus-veneris (Southern Maidenhair Fern) · A. capillus-veneris 'Banksianum' · A. capillus-veneris 'Cornubiense' · A. capillus-veneris 'Fimbriatum' · A. capillus-veneris 'Imbricatum' (Imbricatum Southern Maidenhair Fern Adiantum Capillus-Veneris) · A. capillus-veneris L. 'Fimbriatum' (Fimbriatum Southern Maidenhair Fern Adiantum Capillus-Veneris) · A. capillus-veneris 'Mairisii' · A. capillus-veneris 'Pointonii' · A. capillus-veneris var. fissum · A. capillus-veneris var. trifidum · A. capillus-venerus · A. capillusveneris · A. capillus gorgonis · A. capilus-veneris · A. capitus-junosis

More Info

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Further Reading

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  • Fernald, M. L. 1950b. Adiantum capillus-veneris in the United States. Rhodora 52: 201--208.
  • Paris, C. A. 1991. Adiantum viridimontanum, a new maidenhair fern in eastern North America. Rhodora 93: 105--122.
  • Paris, C. A. and M. D. Windham. 1988. A biosystematic investigation of the Adiantum pedatum complex in eastern North America. Syst. Bot. 13: 240--255.
  • Wagner, W. H. Jr. 1956. A natural hybrid, × Adiantum tracyi C. C. Hall. Madroño 13: 195--205.
  • Notes

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    Contributors

    Data Sources

    Accessed through GBIF Data Portal November 12, 2007:

    Identifiers

    Footnotes

    1. Cathy A. Paris "Adiantum". in Flora of North America Vol. 2. Oxford University Press. Online at EFloras.org. [back]
    2. "Adiantum capillus-veneris". in Flora of North America Vol. 2. Oxford University Press. Online at EFloras.org. [back]
    Last Revised: 2009-08-06