Overview
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Vulnerable |
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Interesting Facts
Description
Family Thymelaeaceae
Shrubs
or small trees
, rarely herbs, evergreen
or deciduous. Bark
tough and fibrous
. Leaves opposite or alternate, rarely some ternate
, estipulate; blade
simple
, entire, pinnately veined, articulate
at base
. Plants
mostly bisexual
, sometimes dioecious. Inflorescences terminal
or subterminal
, less often axillary
, sometimes on brachyblasts
, sessile or pedunculate
, basically racemose, sometimes capitate, spicate
, umbelliform, or fascicled. Flowers usually actinomorphic
, bisexual or unisexual
(plants then mostly dioecious), bracteate
(sometimes bracts forming an involucre) or ebracteate
, sessile or pedicellate
. Calyx tubular
, campanulate
, or infundibuliform
, usually corollalike, 4- or 5(or 6) -merous, mostly caducous
, sometimes circumscissile, or persistent
; lobes
imbricate. Petals absent or represented by 4-12 scales
, inserted
at or near throat
of calyx tube (Aquilaria) . Stamens 2 to many, usually as many as calyx lobes and opposite them or twice as many. Hypognous disk usually present at base of ovary, scalelike, annular
or cup-shaped, sometimes absent. Ovary superior, 1- or 2-loculed, sessile or shortly stipitate
; ovules solitary in each locule, pendulous, anatropous
; style filiform
, caducous, sometimes very short or obscure
, terminal or eccentric
; stigma capitate, globose
, subglobose, subclavate, or pyramidal
, sometimes papilose. Fruit mostly indehiscent, dry or fleshy
, sometimes a loculicidal capsule (Aquilaria) . Seeds with or without endosperm, embryo straight.
About 48 genera and ca.
650 species: widely distributed in both hemispheres; nine genera and 115 species (89 endemic) in China.
The phloem contains very strong
fibers, which make the bark of many species very suitable for the manufacture of high-quality paper such as that used for bank notes
. The stems are extremely supple and difficult to break
and are used as a substitute for string. Most species are poisonous and some are important medicinally.[1]
Genus Aquilaria
Trees
or treelets. Leaves alternate, leaf blade
pinnately veined; veins slender, ascending
toward margins
and joining several intramarginal veins
. Inflorescences axillary
or terminal
, umbelliform or paniculiform
, usually without involucre; peduncle short or absent. Flowers bisexual
, pedicellate
, usually 5-merous. Calyx tube
persistent
, yellow or yellowish green, campanulate
; lobes
5, reflexed
or erect
. Petaloid
appendages
twice as many as the lobes, inserted
at throat
of tube, scalelike, united
in a ring
at base
, usually densely pubescent
or puberulous
. Stamens twice as many as calyx lobes, alternating with petaloid appendages; filaments
short or absent; anthers
oblong
, dorsifixed
, pubescent, connectives
broad. Disk absent. Ovary subsessile
, pubescent, 2-loculed or incompletely 2-loculed; style very short; stigma capitate. Fruit a 2-valved loculicidal capsule, compressed
laterally, obovoid
, base enclosed by persistent calyx; pericarp leathery or woody. Seeds 2 or 1 by abortion
, ovoid
or ellipsoid
, on carunclelike strongly tapered funicle
; testa crustaceous
, endosperm absent; cotyledons thick, plano-convex
.
About 15 species: Bhutan, Cambodia, China, NE India, Laos, Malaysia, Myanmar, Thailand, Vietnam; two species (both endemic) in China.
Members
of this genus are important as the source of agarwood, a very highly prized aromatic
gum produced
in the wood
in response to injury. It is used as incense, in perfumery, and in traditional medicine. Trees have been over-harvested throughout the range
of the genus, and most species are of conservation
concern. The genus as a whole is included
in Category
2 of CITES
.[2]
Habitat
Biome: Terrestrial [3].
Ecology: A small tree occurring in lowland forest on hill slopes .[3].
Taxonomy
- Domain:
Eukaryota
(
)
- Whittaker & Margulis,1978
- eukaryotes
- Kingdom:
Plantae
(
)
- Haeckel, 1866
- Plants
- Subkingdom:
Viridaeplantae
(
)
- Cavalier-Smith, 1981
- Phylum:
Tracheophyta
(
)
- Sinnott, 1935 ex Cavalier-Smith, 1998
- Vascular Plants
- Subphylum:
Euphyllophytina
(
)
- Infraphylum:
Radiatopses
(
)
- Kenrick & Crane, 1997
- Class:
Magnoliopsida
(
)
- Brongniart, 1843
- Dicotyledons
- Subclass:
Rosidae
(
)
- Takhtajan, 1967
- Superorder:
Euphorbianae
(
)
- Takhtajan Ex Reveal, 1992
- Order:
Myrtales
(
)
- Dumortier, 1829
- Family:
Thymelaeaceae
(
)
- A.L. de Jussieu, 1789, nom. cons.
- mezereum
- Subfamily:
Thymelaeoideae
(
)
- Tribe:
Aquilarieae
(
)
- Genus:
Aquilaria
(
)
- Lamarck, 1783, nom. cons.
- Specific epithet:
hirta
- Ridl.
- Botanical name: - Aquilaria hirta Ridl.
- Specific epithet:
hirta
- Ridl.
- Genus:
Aquilaria
(
- Tribe:
Aquilarieae
(
- Subfamily:
Thymelaeoideae
(
- Family:
Thymelaeaceae
(
- Order:
Myrtales
(
- Superorder:
Euphorbianae
(
- Subclass:
Rosidae
(
- Class:
Magnoliopsida
(
- Infraphylum:
Radiatopses
(
- Subphylum:
Euphyllophytina
(
- Phylum:
Tracheophyta
(
- Subkingdom:
Viridaeplantae
(
- Kingdom:
Plantae
(
Notes
Publishing author : Ridl. Publication : J. Straits Branch Roy. Asiat. Soc. 35: 78 1901 [Jan 1901]
Similar Species
Members of the genus Aquilaria
ZipcodeZoo has pages for 2 species, subspecies, varieties, forms, and cultivars in this genus:
A. crassna (Agar Wood) · A. malaccensis (Agarwood)
More Info
- Search for Pictures: images.google.com
- Search for Scholarly Articles: Google Scholar
- Search using Scientific Name and Vernacular Names: All the Web | AltaVista Canada | AltaVista | Excite | Google | HotBot | Lycos
- Search using Specialized Databases: GenBank | Medline | Scirus | CISTI/CAL | Agricola Periodicals | Agricola Books
Further Reading
- Conservation and sustainable management of trees, report of the third regional workshop, held at Army Hotel, Hanoi, Viet Nam, 18-21 August, 1997 WCMC url p. 28.
- Flora Malesiana. general editor, C.G.G.J. van Steenis. Djakarta: Noordhoff-Kolff, 1950- url p. 7.
- The World List of Threatened Trees WCMC, IUCN url p. 61.
- Huang Shuchung & Zhang Zerong. 1999. Thymelaeaceae. In: Ku Tsuechih, ed., Fl. Reipubl. Popularis Sin. 52(1): 287-400.
- Huang Shuchung & Zhang Zerong. 1999. Thymelaeaceae. In: Ku Tsuechih, ed., Fl. Reipubl. Popularis Sin. 52(1): 287-400.
Notes
Contributors
- Brands, S.J. (comp.) 1989-present. The Taxonomicon. Universal Taxonomic Services, Zwaag, The Netherlands. Accessed January 15, 2012.
- IUCN 2012. IUCN Red List of Threatened Species. Version 2011.2. . Downloaded on January 28, 2012.
- World Conservation Monitoring Centre 1998. Aquilaria hirta. In: IUCN 2011. IUCN Red List of Threatened Species. Version 2011.2. <www.iucnredlist.org>. Downloadedon 30January2012.
Identifiers
- Biodiversity Heritage Library NamebankID: 5924252
- Global Biodiversity Information Facility Taxonkey: 15880519
- Globally Unique Identifier: urn:lsid:ipni.org:names:20001505-1
- International Plant Names Index (IPNI) ID: 20001505
- IUCN ID: 192283
- Zipcode Zoo Species Identifier: 1022330
Footnotes
- Yinzheng Wang, Michael G. Gilbert, Brian F. Mathew, Christopher Brickell & Lorin I. Nevling "Thymelaeaceae". in Flora of China Vol. 13 Page 213. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org. [back]
- Yinzheng Wang, Lorin I. Nevling & Michael G. Gilbert "Aquilaria". in Flora of China Vol. 13 Page 213, 214. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org. [back]
- World Conservation Monitoring Centre 1998. Aquilaria hirta. In: IUCN 2011. IUCN Red List of Threatened Species. Version 2011.2. <www.iucnredlist.org>. Downloaded on 30 January 2012. [back]
