Overview
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Extirpated |
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Common Names
Common Names in English:
Crenate-Lobed False Goat's-Beard, Roan False Goats Beard, Roan Mountain False Goat's Beard
Description
Family Saxifragaceae
Herbs or shrubs
, rarely trees
or vines
. Leaves simple
or compound
, usually alternate or opposite, usually exstipulate
. Flowers usually in cymes, panicles, or racemes
, rarely solitary, usually bisexual
, rarely unisexual
, hypogynous or ± epigynous
, rarely perigynous, usually biperianthial, rarely monochlamydeous
, actinomorphic
, rarely zygomorphic, 4- or 5(-10) -merous. Sepals sometimes petal-like. Petals usually free
, sometimes absent. Stamens (4 or) 5-10 or many; filaments
free; anthers
2-loculed; staminodes often present. Carpels 2, rarely 3-5(-10), usually ± connate
; ovary superior or semi-inferior to inferior, 2- or 3-5(-10) -loculed with axile placentation
, or 1-loculed with parietal placentation
, rarely with apical placentation; ovules usually many, 2- to many seriate
, crassinucellate
or tenuinucellate
, sometimes with transitional forms; integument 1- or 2-seriate; styles free or ± connate. Fruit a capsule or berry, rarely a follicle or drupe. Seeds albuminous
, rarely not so; albumen of cellular
type, rarely of nuclear
type; embryo small.
About 80 genera and 1200 species: worldwide; 29 genera (two endemic), and 545 species (354 endemic, seven introduced
) in China.
During the past several years, cladistic analyses of morphological, chemical, and DNA data have made it clear that the recognition of the Saxifragaceae sensu
lato (Engler, Nat. Pflanzenfam. 18a: 74-226. 1928) is untenable. Among the angiosperm families, Saxifragaceae sensu lato may in fact represent the most extreme example of a polyphyletic assemblage
. For example, recent analyses of DNA sequence data indicate that these taxa represent at least ten separate evolutionary lines
, many of which are only distantly related to one another (Morgan & Soltis, Ann. Missouri Bot. Gard. 80: 631-660. 1993; Soltis & Soltis, Amer. J. Bot. 84: 504-522. 1997) . Furthermore, very large molecular phylogenetic
analyses of hundreds
of angiosperms
indicate that these separate lineages
are distributed among four of the six traditionally recognized subclasses of dicotyledons (Savolainen et al.
, Syst. Biol. 49: 306-362. 2000; Soltis et al., Nature 402: 402-404. 1999) . These recent studies have also greatly clarified how this phylogenetically diverse
assemblage should be divided
into families and treated taxonomically (see The Angiosperm Phylogeny Group (APG), Ann. Missouri Bot. Gard. 85: 531-553. 1998) . Recent studies of DNA sequence data have clarified both the circumscription and affinities of a narrowly defined Saxifragaceae (Saxifragaceae sensu stricto) and Hydrangeaceae (Soltis et al., Amer. J. Bot. 82: 504-514. 1995; Savolainen et al., loc. cit.
; Soltis et al., loc. cit. 1999) . Saxifragaceae sensu stricto should consist only of Saxifragoideae, a group of about 30 herbaceous genera. Members
of Saxifragaceae sensu stricto from the Chinese flora
include Astilbe, Astilboides, Bergenia, Chrysosplenium, Mitella, Mukdenia, Oresitrophe, Rodgersia, Saxifraga, Tanakaea, Tiarella, and the recently described Saniculiphyllum. Close relatives of Saxifragaceae sensu stricto include Itea, Penthorum, and Ribes. These genera, the sole
members of Iteoideae, Penthoroideae, and Ribesioideae, respectively, are also best treated in separate families: Iteaceae, Penthoraceae, and Grossulariaceae (see APG, loc. cit.) . These taxa, as well as several others, such as Crassulaceae, are basal to a large assemblage of taxa, most of which were traditionally placed in Rosidae
. Sequence data also indicate that Parnassia (the sole member of the Parnassioideae) is a more derived member of the rosid alliance
, most closely related to Brexia and Lepuropetalon (also part of Saxifragaceae sensu lato) and Celastraceae. Parnassia and Lepuropetalon should be placed in Parnassiaceae with Brexia part of an expanded Celastraceae (APG, loc. cit.) .
Both morphological and molecular data indicate that Hydrangeoideae and Escallonioideae are, in contrast, allied with taxa traditionally placed in Asteridae. Hydrangeoideae are a well-defined, monophyletic lineage that should be treated as Hydrangeaceae. In China they include Cardiandra, Decumaria, Deinanthe, Deutzia, Dichroa, Hydrangea, Kirengeshoma, Philadelphus, Pileostegia, Platycrater, and Schizophragma, and are closely allied with families such as Cornaceae, Loasaceae, and Nyssaceae. Escallonioideae appear to be polyphyletic, and this group of approximately 14 genera is in need of thorough study. Members of this subfamily
are allied with several different lineages of higher asterids. Polyosma, the only member of Escallonioideae in China, appears closely allied with Caprifoliaceae (Xiang & Soltis in Boufford & Ohba, Sino-Japanese Flora: its Characteristics and Diversification, 1998) .[1]
Genus Astilbe
erbs perennial
. Rhizomes thick. Stems brown paleaceous
hairy
or long pilose
. Leaves alternate, long petiolate
, 2-4 × ternately compound
, rarely simple
; stipules membranous; leaflets
lanceolate, ovate
, or broadly ovate to elliptic
, margin
dentate
. Inflorescence a terminal
panicle, bracteate
. Flowers white, lilac, or purple, bisexual
or unisexual
, rarely plants
polygamous or dioecious. Sepals (4 or) 5. Petals usually 1-5, sometimes more or absent. Stamens usually (5 or) 8-10. Carpels 2(or 3), ± connate
or free
; ovary subsuperior or semi-inferior, 2(or 3) -loculed with axile placentation
or 1-loculed with marginal placentation
; ovules many. Fruit a capsule or follicle. Seeds small.
About 18 species: Asia, North America: seven species (three endemic) in China.[2]
Physical Description
Habit: Forb/herb
Foliage: Summer foliage: Serrated compound leaves attractive all season .
Size/Age/Growth
Size: 22-26" tall.
Biology
Reproduction
Duration: Perennial
Growth
Culture: Space 18-24" apart.
Soil: Rich, humusy, moist soil. Needs moist soil.
Sunlight: Sun Exposure: Full Sun .
Moisture: Water Requirements: Keep soil moist.
Temperature: Cold Hardiness: 3a, 3b, 4a, 4b, 5a, 5b, 6a, 6b, 7a, 7b, 8a, 8b, 9a, 9b. (map)
Taxonomy
- Domain:
Eukaryota
(
)
- Whittaker & Margulis,1978
- eukaryotes
- Kingdom:
Plantae
(
)
- Haeckel, 1866
- Plants
- Subkingdom:
Viridaeplantae
(
)
- Cavalier-Smith, 1981
- Phylum:
Tracheophyta
(
)
- Sinnott, 1935 Ex Cavalier-Smith, 1998
- Vascular Plants
- Subphylum:
Euphyllophytina
(
)
- Infraphylum:
Radiatopses
(
)
- Kenrick & Crane, 1997
- Class:
Magnoliopsida
(
)
- Brongniart, 1843
- Dicotyledons
- Subclass:
Rosidae
(
)
- Takhtajan, 1967
- Superorder:
Saxifraganae
(
)
- Reveal, 1994
- Order:
Saxifragales
(
)
- Dumortier, 1829
- Family:
Saxifragaceae
(
)
- Durande, 1782, Nom. Cons.
- Saxifrage Family
- Genus:
Astilbe
(
)
- Buchanan-Hamilton ex D. Don, Prodr. Fl. Nepal. 210. 1825.
- False Goat's Beard
- Specific epithet:
crenatiloba
- Small
- Botanical name: - Astilbe crenatiloba
- Specific epithet:
crenatiloba
- Small
- Genus:
Astilbe
(
- Family:
Saxifragaceae
(
- Order:
Saxifragales
(
- Superorder:
Saxifraganae
(
- Subclass:
Rosidae
(
- Class:
Magnoliopsida
(
- Infraphylum:
Radiatopses
(
- Subphylum:
Euphyllophytina
(
- Phylum:
Tracheophyta
(
- Subkingdom:
Viridaeplantae
(
- Kingdom:
Plantae
(
Notes
Name Status: Accepted Name . Latest taxonomic scrutiny: 15-Mar-2000
Similar Species
Members of the genus Astilbe
ZipcodeZoo has pages for 437 species, subspecies, varieties, forms, and cultivars in this genus. Here are just 100 of them:
A. 'Alive and Kicking' · A. 'Amerika' · A. 'Amethyst' · A. 'Angel Wings' · A. 'Anita Pfeifer' · A. 'Aphrodite' · A. apoensis · A. ardensii-Hybride · A. arendsii · A. x arendsii 'Flamingo' (Flamingo Astilbe) · A. arendsi Diamond · A. aruncus · A. astilboides · A. 'Atrorosea' · A. 'Augustleuchten' (August Light False Spirea) · A. austrosinensis · A. 'Avalanche' · A. bandaica · A. barbata · A. Bella Group · A. 'Bergkristall' · A. 'Betsy Cuperus' · A. biternata (Appalachian False Goat's Beard) · A. 'Bonn' · A. 'Brautschleier' · A. 'Bremen' · A. Bridal Veil · A. 'Broncelaub' · A. 'Bronce Elegans' · A. 'Bronze Sprite' · A. 'Brunhilde' · A. 'Bumalda' · A. bumalda 'Bronze Pygmy' · A. 'Burgunderrot' · A. 'Carminea' · A. 'Carmine King' · A. 'Carnea' · A. 'Catherine Deneuve' · A. 'Cattleya' · A. 'Cattleya Dunkel' · A. 'Ceres' · A. 'Cherry Ripe' · A. chinensis (False Goats Beard) · A. chinensis 'Alba' · A. chinensis 'Brokat' · A. chinensis (Maxim.) Franch. & Sav. var. chinensis · A. chinensis 'Christian' · A. chinensis 'Crispa Perkeo' (False Goats-Beard) · A. chinensis 'Finale' (False Spirea) · A. chinensis 'Frankentroll' · A. chinensis 'Intermezzo' (False Goats-Beard) · A. chinensis 'Maggie Daley' (Maggie Daley False Goats-Beard) · A. chinensis 'Milk and Honey' · A. chinensis 'Pumila' · A. chinensis (Maxim.) Franch. & Sav. var. pumila hort. 'Liliput' · A. chinensis 'Purple Glory' · A. chinensis 'Purple Lance' · A. chinensis 'Rise and Shine' · A. chinensis 'Superba' · A. chinensis (Maxim.) Franch. & Sav. 'Superba' (False Goats Beard) · A. chinensis 'Tacquettii Superba' · A. chinensis (Maxim.) Franch. & Sav. var. taquetii · A. chinensis (Maxim.) Franch. & Sav. var. taquetii 'Purpurlanze' · A. chinensis (Maxim.) Franch. & Sav. var. taquetii 'Superba' · A. chinensis 'Troll' · A. chinensis 'Valery' · A. chinensis var. davidii (False Goats-Beard) · A. chinensis var. pumila (False Goats Beard) · A. chinensis var. pumila 'Serenade' · A. chinensis var. taquetii (False Goats-Beard) · A. chinensis var. taquetii 'Purpurlanze' · A. chinensis var. taquetii 'Red Charm' (False Goats-Beard) · A. chinensis var. taquetii 'Rowden Sunstar' · A. chinensis var. taquetii 'Superba' · A. chinensis var. taquetii 'Susannah' · A. chinensis var. taquetii 'Purple Candles' · A. chinensis 'Veronica Klose' (False Goats Beard) · A. chinensis 'Veronika Klose' · A. chinensis 'Visions' (Visions False Goats Beard) · A. chinensis 'Visions in Pink' (False Goats Beard) · A. chinensis 'Visions in Red' (Visions in Red False Goats Beard) · A. chinensis 'Vision in Pink' · A. chinensis white · A. 'Color Flash' · A. 'Color Flash Lime' · A. congesta · A. congesta var. bandaica · A. 'Country and Western' · A. crenatiloba (Crenate-Lobed False Goat's-Beard) · A. crenatilobata · A. 'Crimson Feather' · A. crispa · A. 'Darwin's Dream' · A. 'Darwin's Favourite' · A. 'Darwin's Surprise' · A. davidii · A. decandra · A. decandra crenatiloba · A. decandra var. crenatiloba · A. 'Delft Lace'
More Info
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Further Reading
- Botanisches Zentralblatt; referierendes Organ für das Gesamtgebiet der Botanik. Jena [etc.]G. Fischer [etc.] GER url p. 380.
- Hwang Shu-mei, Wei Chao-fen, Lu Ling-ti, Ku Tsue-chih &
- Hwang Shu-mei, eds., Fl. Reipubl. Popularis Sin. 35(1): 1-406.
- Jin Shu-ying. 1995. Saxifragaceae (2) [Parnassioideae, Hydrangeoideae, Escallonioideae, Iteoideae, Ribesioideae]. In: Lu Ling-ti &
- Pan Jin-tang. 1992. Saxifragaceae (1) [Penthoroideae, Saxifragoideae]. In: Pan Jin-tang, ed., Fl. Reipubl. Popularis Sin. 34(2): 1-309
- Hwang Shu-mei, Wei Chao-fen, Lu Ling-ti, Ku Tsue-chih & Jin Shu-ying. 1995. Saxifragaceae (2) [Parnassioideae, Hydrangeoideae, Escallonioideae, Iteoideae, Ribesioideae]. In: Lu Ling-ti & Hwang Shu-mei, eds., Fl. Reipubl. Popularis Sin. 35(1): 1-406.
- Pan Jin-tang. 1992. Saxifragaceae (1) [Penthoroideae, Saxifragoideae]. In: Pan Jin-tang, ed., Fl. Reipubl. Popularis Sin. 34(2): 1-309
Notes
Contributors
- Global Biodiversity Information Facility. Accessed February 02, 2008. http://www.gbif.org Mediated distribution data from 2 providers.
Data Sources
Accessed through GBIF Data Portal February 02, 2008:
- USDA PLANTS, USDA PLANTS Database
Identifiers
- Biodiversity Heritage Library NamebankID: 2649625
- Catalogue of Life Accepted Name Code: ITS-24306
- Global Biodiversity Information Facility Taxonkey: 13742857
- Globally Unique Identifier: urn:lsid:ipni.org:names:790274-1
- Integrated Taxonomic Information System (ITIS) Taxonomic Serial Number (TSN): 24306
- International Plant Names Index (IPNI) ID: 24376-2
- Natural Heritage Network Species Identifier: PDSAX01020
- U.S.D.A. Plant Symbol: ASCR7
- Zipcode Zoo Species Identifier: 21169
Footnotes
- Jin-tang Pan, Cuizhi Gu, Shumei Huang, Chao-fen Wei, Shu-ying Jin, Lingdi Lu, Shinobu Akiyama, Crinan Alexander, Bruce Bartholomew, James Cullen, Richard J. Gornall, Ulla-Maj Hultgård, Hideaki Ohba & Douglas E. Soltis "Saxifragaceae". in Flora of China Vol. 8 Page 269. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org. [back]
- Pan Jintang , Hideaki Ohba "Astilbe". in Flora of China Vol. 8 Page 274. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org. [back]
