Overview
Interesting Facts
Common Names
Click on the language to view common names.
Common Names in English:
Cockspur Thistle, Malta Starthistle, Malta Thistle, Maltese Star Centaury, Maltese Star Thistle, Maltese Star-Thistle, Maltese Starthistle, Napa Thistle, Spotted Knapweed, Tocalote, Yellow Star-Thistle
Common Names in French:
Coix De Malte
Common Names in German:
Malteser Flockenblume
Common Names in Portuguese:
Centáurea-Estrela-De-Malta
Description
Family Compositae
The largest family of flowering plants , the Compositae (Asteraceae), comprising about 1,100 genera and more than 20,000 species and characterized by many small flowers arranged in a head looking like a single flower and subtended by an involucre of bracts. A head may consist of both ray flowers and disk flowers, as in the sunflower, of disk flowers only, as in the burdock, or of ray flowers only, as in the dandelion.
Genus Centaurea
Annuals
, biennials, or perennials, 20-300 cm, glabrous
or tomentose
. Stems erect
, ascending
, or spreading
, simple
or branched. Leaves basal and cauline; petiolate
or sessile; proximal
blade
margins
often ± deeply lobed
, (spiny
in C. benedicta ), distal ± smaller, often entire, faces
glabrous or ± tomentose, sometimes also villous
, strigose
, or puberulent
, often glandular-punctate. Heads discoid
, disciform
, or radiant, borne singly or in corymbiform
arrays. Involucres cylindric
or ovoid
to hemispheric
. Phyllaries many in 6-many series, unequal, proximal part appressed
, body margins entire. distal parts expanded into erect to spreading, usually ± dentate
or fringed
, linear
to ovate
appendages
, spine. tipped or spineless. Receptacles flat, epaleate, bristly
. Florets 10-many; outer usually sterile
, corollas slender and inconspicuous to much expanded, ± bilateral
; inner fertile
, corollas white to blue, pink, purple, or yellow, bilateral or radial
, often bent at junction of tubes
and throats
, lobes
linear-oblong, acute; anther
bases
tailed
, apical appendages oblong
; style branches: fused portions with minutely hairy
nodes, distinct
portions minute. Cypselae ± barrel-shaped, ± compressed
, smooth
or ribbed
, apices entire (denticulate
in C.
benedicta ), glabrous or with fine, 1-celled hairs
, attachment scar
. lateral
(with or without elaiosomes) ; pappi 0 or ± persistent
, of 1-3 series of smooth or minutely barbed
, stiff bristles
or narrow scales
. x = 8, 9, 10, 11, 12, 13, 15.
Species ca.
500: introduced
; Eurasia
, n Africa, widely introduced worldwide.
Taxonomic
limits
of Centaurea have been controversial. The genus has great morphologic diversity
, and studies have revealed much cytologic (e.g.
, N. Garcia-Jacas et al.
1996) and palynologic (e.g., G. Wagenitz 1955) variation
as well. During the nineteenth and twentieth centuries, various taxonomists attempted, with limited success, to divide Centaurea into smaller genera or workable infrageneric
taxa. The relations of several satellite
genera have been controversial as well.
Recent molecular phylogenetic
studies (A. Susanna et al. 1995; N. Garcia-Jacas et al. 2000, 2001) have begun to clarify relationships
within Centaurea and between Centaurea and other genera. These studies make it clear that Centaurea as traditionally defined is polyphyletic, and that generic
boundaries should be realigned if monophyletic taxa are to be recognized. Some taxa traditionally included
within Centaurea (e.g., the two native
North American species, Centaurea americana and C. rothrockii) fall
outside the redefined generic boundaries and are here treated in Plectocephalus. Others usually placed into segregate
genera (e.g., Cnicus benedictus) are firmly nested within Centaurea. Because the type species of Centaurea (C. centaurium Linnaeus, an African species) falls
outside the main lineage
of the genus, a proposal
has been made to conserve Centaurea with a different type species (W. Greuter et al. 2001), thereby maintaining the nomenclatural
stability
of most of the numerous
species that do fall within the principal Centaurea clade.
Although several Centaurea species are widely established
as members
of the North American flora
, and some of these are widely distributed invasive weeds
, some of the taxa listed by J. T. Kartesz and C. A. Meacham (1999) are apparently waifs
and not permanent members of the flora. These taxa are discussed informally immediately below.
Although Cnicus has usually been recognized as a distinctive monotypic genus, it has been merged into Centaurea by various authors
(e.g., K
. Bremer 1994; G. Wagenitz and F. H. Hellwig 1996) . Recent molecular systematic studies (N. Garcia-Jacas et al. 2000) provide additional evidence that it is nested within Centaurea.[1]
Physical Description
Species Centaurea melitensis
Annuals, 10-100 cm, herbage
loosely gray-tomentose and villous
with jointed
multicellular
hairs
, sometimes minutely scabrous
, minutely
resin-gland-dotted. Stems 1-few, few-many branched distally.
Leaves: basal and proximal
cauline petiolate
or tapering to
base
, usually absent at anthesis
, blades
oblong
to oblanceolate
,
2-15 cm, margins
entire to dentate
or pinnately lobed
; cauline long-decurrent,
blades linear
to oblong or oblanceolate, 1-5 cm, entire or dentate.
Heads disciform
, 1-few at branch
tips
, borne singly or in
open leafy corymbiform
arrays, sometimes clustered in distal axils,
sessile or pedunculate
. Involucres ovoid
, 10-15 mm, loosely
cobwebby-tomentose or becoming glabrous
. Principal phyllaries:
bodies ± stramineous
, ovate
, appendages
purplish, spiny-fringed
at base, each tipped by slender spine 5-10 mm.
Inner phyllaries:
appendages entire, acute or spine-tipped. Florets many; corollas
yellow, those of sterile
florets
10-12 mm, slender, inconspicuous,
those of fertile
florets 10-12 mm. Cypselae dull
white or
light brown, ca.
2.5 mm, finely hairy
; pappi of many white,
unequal, stiff bristles
2.5-3 mm. 2n = 24. Flowering
mostly spring-summer (Apr-Jul). [source]
Centaurea melitensis is native
to the Mediterranean region.
It is listed as a noxious weed
in New Mexico. [source]
Habit: Forb/herb
Flowers: Bloom Period: March, April, May, June. • Flower Color: yellow
Size/Age/Growth
Size: 12-18" tall.
Habitat
Roadsides, fields , pine-oak woodlands, chaparral , agricultural areas; 0-1500 m ; widely introduced [2].
Typically found at an altitude of 0 to 2,147 meters (0 to 7,044 feet).[3]
Ecology: ReproductionDiTomaso (2001) reports that C. melitensis is insect-pollinated and reproduces by seed. Seed production is highly variable. Plants can produce 1-60 or more seeds per head and 1-100 heads or more per plant.
Biology
Reproduction
Duration: Annual , Biennial
Growth
Soil: Minimum pH: 6.1 • Maximum pH: 8.5
Sunlight: Sun Exposure: Full Sun .
Taxonomy
- Domain:
Eukaryota
(
)
- Whittaker & Margulis,1978
- eukaryotes
- Kingdom:
Plantae
(
)
- Haeckel, 1866
- Plants
- Subkingdom:
Viridaeplantae
(
)
- Cavalier-Smith, 1981
- Phylum:
Tracheophyta
(
)
- Sinnott, 1935 ex Cavalier-Smith, 1998
- Vascular Plants
- Subphylum:
Euphyllophytina
(
)
- Class:
Spermatopsida
(
)
- Brongniart, 1843
- Subclass:
Asteridae
(
)
- Takhtajan, 1967
- Superorder:
Campanulanae
(
)
- Takhtajan Ex Reveal, 1992
- Order:
Asterales
(
)
- Lindley, 1833
- Family:
Compositae
(
)
- Giseke, 1792, nom. cons., nom. alt.
- Subfamily:
Carduoideae
(
)
- Tribe:
Cardueae
(
)
- Subtribe:
Centaureinae
(
)
- Genus:
Centaurea
(
)
- C. Linnaeus, 1753
- Knapweed, star thistle, cornflower [Greek kentaurieon, ancient plant name associated with Chiron, a centaur famous for knowledge of medicinal plants]
- Specific epithet:
melitensis
- L.
- Botanical name: - Centaurea melitensis L.
- Specific epithet:
melitensis
- L.
- Genus:
Centaurea
(
- Subtribe:
Centaureinae
(
- Tribe:
Cardueae
(
- Subfamily:
Carduoideae
(
- Family:
Compositae
(
- Order:
Asterales
(
- Superorder:
Campanulanae
(
- Subclass:
Asteridae
(
- Class:
Spermatopsida
(
- Subphylum:
Euphyllophytina
(
- Phylum:
Tracheophyta
(
- Subkingdom:
Viridaeplantae
(
- Kingdom:
Plantae
(
Notes
Name
Status: Accepted Name
.
Comment: Data Providers: CONABIO, African Flowering Plants
Database
,
SANBI, New Zealand Plant Name Database, Govaerts World Compositae
Checklist
A-G, IPNI, Tropicos, Euro+Med, Flora
of Tasmania Online,
Northern Territory Checklist, Queensland Census, New South Wa. GCC
LSID: urn
:lsid:compositae.org:names:A83F8D03-79DA-4938-8C82-A516DEB694A6
Last scrutiny: 14-Nov-09
Similar Species
Members of the genus Centaurea
ZipcodeZoo has pages for 82 species, subspecies, varieties, forms, and cultivars in this genus:
C. akamantis (Akamas Centaury) · C. alpestris (Greater Centaury) · C. americana (American Basketflower) · C. americana 'Aloha' (American Knapweed) · C. americana 'Jolly Joker' (American Knapweed) · C. aspera (Rough Star-Thistle) · C. atropurpurea (Centaurea) · C. bella (Knapweed) · C. benoistii (Maroon Cornflower) · C. biebersteinii (Ballast Waif Centaurea Biebersteinii) · C. bovina (Pasture Knapweed) · C. cachinalensis (Flor Del Minero) · C. calcitrapa (Big-Head Purple Starthistle) · C. calcitrapoides (Smallhead Star-Thistle) · C. cineraria (Dusty Miller) · C. cineraria 'Colchester White' (Dusty Miller) · C. crupina (Crupina) · C. cyanoides 'Blue Carpet' (Cornflower) · C. cyanus (Bachelor's Button) · C. cyanus nana 'Jubilee Gem' (Bachelors Button) · C. cyanus 'Black Ball' (Bachelors Button) · C. cyanus 'Black Boy' (Bachelors Button) · C. cyanus 'Black Gem' (Bachelors Button) · C. cyanus 'Black Magic' (Bachelors Button) · C. cyanus 'Blue Boy' (Bachelors Button) · C. cyanus 'Blue Diadem' (Bachelors Button) · C. cyanus 'Classic Blue' (Bachelors Button) · C. cyanus 'Dwarf Blue Midget' (Bachelors Button) · C. cyanus 'Frosted Queen Mix' (Bachelors Button) · C. cyanus 'Frosty Mix' (Bachelors Button) · C. cyanus 'Mauve Queen' (Bachelors Button) · C. cyanus 'Polka Dot Mix' (Bachelors Button) · C. dealbata (Knapweed Centaurea Dealbata) · C. dealbata 'Rosea' (Knapweed) · C. debeauxii (Meadow Knapweed) · C. debeauxii thuillieri (Meadow Knapweed) · C. debeauxii subsp. thuillieri (Meadow Knapweed) · C. depressa (Centaurea) · C. diffusa (Diffuse Knapweed) · C. diluta (North African Knapweed) · C. eriophora (Sand-Heath) · C. gymnocarpa (Velvet Centaurea) · C. hypoleuca DC. 'John Coutts' (Knapweed) · C. iberica (Iberian Knapweed) · C. jacea (Brown Knapweed) · C. jacea x nigra (Hybrid Knapweed Centaurea Jacea X Nigra) · C. macrocephala (Armenian Basketflower) · C. melitensis (Cockspur Thistle) · C. moncktonii (Meadow Knapweed) · C. montana (Mountain Bluet) · C. montana 'Alba' (Mountain Bluet) · C. montana 'Amethyst in Snow' (Mountain Bluet) · C. montana 'Dot Purple' (Mountain Bluet) · C. montana 'Gold Bullion' (Batchelor's Button) · C. moschata 'Dairy Maid' (Sweet Sultan) · C. moschata 'Imperialis' (Sweet Sultan) · C. moschata 'The Bride' (Sweet Sultan) · C. nigra (Black Knapweed) · C. nigrescens (Short-Fringe Starthistle) · C. orientalis (Centaurea) · C. ovina (Lilac Knapweed) · C. paniculata (Jersey Knapweed) · C. phrygia (Scandinavian Starthistle) · C. pindicola (Centaurea) · C. pulcherrima (Pink Bachelors Button) · C. 'Pulchra Major' (Bachelor's Button) · C. ragusina (Dubrovacka Zecina) · C. rothrockii (Basket Flower) · C. scabiosa (Great Starthistle) · C. solstitialis (Barnaby Star-Thistle) · C. solstitialis solstitialis (St. Barnaby's Thistle) · C. stoebe (Spotted Knapweed) · C. sulphurea (Sicilian Starthistle) · C. thuillieri (Meadow Knapweed) · C. transalpina (Alpine Knapweed) · C. trichocephala (Feather-Head Knapweed) · C. triumfetti (Squarrose Knapweed Centaurea Triumfetti) · C. triumfettii (Spreading Thistle) · C. uniflora (Single-Flower Knapweed) · C. uniflora nervosa (Singleflower Knapweed) · C. xpratensis (Meadow Knapweed) · C.'Nigra' (Bachelor Buttons)
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Further Reading
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Notes
Contributors
- Brands, S.J. (comp.) 1989-present. The Taxonomicon. Universal Taxonomic Services, Zwaag, The Netherlands. Accessed March 27, 2012.
Data Sources
Accessed through GBIF Data Portal November 17, 2007:
- Australian National Herbarium
- , Australian National Herbarium
- Berkeley Natural History Museums, University and Jepson Herbaria DiGIR provider
- Bernice Pauahi Bishop Museum, Bishop Museum Natural History Specimen Data
- Biologiezentrum der Oberoesterreichischen Landesmuseen, Biologiezentrum Linz
- Bundesamt für Naturschutz / Zentralstelle für Phytodiversität Deutschland, Bundesamt fuer Naturschutz / Zentralstelle fuer Phytodiversitaet Deutschland
- Conservatoire botanique national du Bassin parisien, Conservatoire botanique national du Bassin parisien
- European Environment Agency, EUNIS
- GBIF-Spain, BDBCV BioBlitz in Penyagolosa
- GBIF-Spain, Botánica, Universidad de León: LEB-Cormo
- GBIF-Spain, Dirección General de Investigación, Desarrollo Tecnológico e Innovación de la Junta de Extremadura(DGIDTI): HSS
- GBIF-Spain, Herbario Universidad de Málaga: MGC-Cormófitos
- GBIF-Spain, Herbario de la Universidad de Sevilla, SEV
- GBIF-Spain, Hortus Botanicus Sollerensis Herbarium
- GBIF-Spain, Jardín Botánico de Córdoba: Herbarium COA
- GBIF-Spain, Real Jardin Botanico
- , Vascular Plant Herbarium
- GBIF-Spain, Universidad de Almería, HUAL
- Missouri Botanical Garden, Missouri Botanical Garden
- National Herbarium of New South Wales, NSW herbarium collection
- Natural History Museum, University of Oslo, Vascular Plant Herbarium, Oslo
- Oregon State University, Vascular Plant Collection
- School of Life Sciences, Arizona State University, Arizona State University Vascular Plant Herbarium
- School of Life Sciences, Arizona State University, The Deaver Herbarium, Northern Arizona University
- SysTax, Herbarium Universitat Ulm
- SysTax, Justus-Liebig-Universitat Giessen
- SysTax, Museu Botanico Municipal
- SysTax, SysTax
- The Swedish Museum of Natural History
- , Botany
- The Swedish Museum of Natural History
- , Herbarium of Oskarshamn
- The Swedish Museum of Natural History
- , Lund Botanical Museum
- UK National Biodiversity Network, Botanical Society of the British Isles - Vascular Plants Database
- USDA PLANTS, USDA PLANTS Database
- University of Alabama Biodiversity and Systematics, Herbarium
- Utah State University, USU-UTC Specimen Database
Identifiers
- Biodiversity Heritage Library NamebankID: 2658263
- Catalogue of Life Accepted Name Code: Ast-596
- Global Biodiversity Information Facility Taxonkey: 4490920
- Globally Unique Identifier: urn:lsid:ipni.org:names:57004-3
- GRIN Nomen Number: 316613
- Integrated Taxonomic Information System (ITIS) Taxonomic Serial Number (TSN): 36965
- Natural Heritage Network Species Identifier: PDAST1Y0D0
- U.S.D.A. Plant Symbol: CEME2
- Zipcode Zoo Species Identifier: 13571
Footnotes
- David J. Keil, Jörg Ochsmann "Centaurea". in Flora of North America Vol. 19, 20 and 21 Page 52, 57, 58, 67, 83, 84, 96, 171, 172, 176, 177, EFloras.org. [back]
- "Centaurea melitensis". in Flora of North America Vol. 19, 20 and 21 Page 183, 193. Oxford University Press. Online at EFloras.org. [back]
- Mean = 370.470 meters (1,215.453 feet), Standard Deviation = 411.860 based on 708 observations. Altitude information for each observation from British Oceanographic Data Centre. [back]
