Description
Family Cycadaceae
Trees
or shrubs
evergreen
, dioecious, mostly palmlike; trunk
columnar
, sometimes dichotomously branched at apex, rarely ovoid-bulbous and subterranean
, clothed with bases
of fallen leaves; bark
often thickened and roughened. Leaves borne at apex of trunk, 1(-3) -pinnately compound
, spirally arranged
; new leaves erect
(or somewhat inflexed
and appearing coiled in C.
multipinnata group), with circinnate leaflets
; petiole
with spines representing (reduced basal leaflets), rarely unarmed
, base swollen and hairy
; leaflets numerous
, alternate to subopposite, dichotomously branched in a few species, midvein
present, margin
usually entire. Cataphylls prominent
, alternate with leaves, hairy, apex often rigid
and pungent
. Pollen cones borne at apex of trunk, cylindric
or fusiform
; microsporophylls
numerous, scalelike, spirally and tightly arranged along axis of cone, with numerous microsporangia in groups abaxially; pollen tubes
producing 2 motile
sperm
cells
. Megasporophylls several to numerous, somewhat leaflike, alternating with flushes of leaves, arranged in a loose
, "conelike" crown surrounding apex of trunk, each with a linear
fertile
stalk
and an apical, pinnatifid
or subentire
sterile
blade
; ovules (1 or) 2-5 on each side of stalk. Seeds drupelike, somewhat compressed
; seed coat
3-layered, consisting of colored
sarcotesta
, woody sclerotesta
, and membranous endotesta
. Cotyledons 2, united
at base. Germination hypogeal, cryptocotylar
. 2n = 22*.
One genus and ca.
60 species: E Africa (including Madagascar), E and S Asia, N Australia, Pacific Islands; 16 species (eight endemic) in China.
Ornamental species
include Cycas
revoluta, which is widely cultivated worldwide. Other species (e.g.
, C. circinalis Linnaeus, C. media R. Brown, C. pectinata, C. rumphii Miquel, C. taitungensis, and C. thouarsii R. Brown) have excellent ornamental
qualities. The stem starch
, "sago" (not to be confused with the true sago as obtained from palms of the genus Metroxylon Rottboøll), is edible and is used in packing brewers’ yeast after the removal of cycasins which are highly toxic
and carcinogenic
. A paste
of Cycas seeds and coconut oil
is used for the treatment of skin
complaints, wounds, ulcers, sores, and boils
.[1]
Genus Cycas
Morphological characters and geographical distribution are the same as those for the family .[2]
Habitat
Typically found at an altitude of 0 to 875 meters (0 to 2,871 feet).[3]
Taxonomy
- Domain:
Eukaryota
(
)
- Whittaker & Margulis,1978
- eukaryotes
- Kingdom:
Plantae
(
)
- Haeckel, 1866
- Plants
- Subkingdom:
Viridaeplantae
(
)
- Cavalier-Smith, 1981
- Phylum:
Tracheophyta
(
)
- Sinnott, 1935 Ex Cavalier-Smith, 1998
- Vascular Plants
- Subphylum:
Euphyllophytina
(
)
- Infraphylum:
Radiatopses
(
)
- Kenrick & Crane, 1997
- Class:
Cycadopsida
(
)
- Order:
Cycadales
(
)
- Dumortier, 1829
- Family:
Cycadaceae
(
)
- Persoon, 1807, Nom. Cons.
- Cycas Family
- Genus:
Cycas
(
)
- C. Linnaeus, 1753
- Cycad
- Specific epithet:
spherica
- Roxb.
- Botanical name: - Cycas spherica Roxb.
- Specific epithet:
spherica
- Roxb.
- Genus:
Cycas
(
- Family:
Cycadaceae
(
- Order:
Cycadales
(
- Class:
Cycadopsida
(
- Infraphylum:
Radiatopses
(
- Subphylum:
Euphyllophytina
(
- Phylum:
Tracheophyta
(
- Subkingdom:
Viridaeplantae
(
- Kingdom:
Plantae
(
Notes
Name Status: Accepted Name . Latest taxonomic scrutiny: Wilson K .L.
Similar Species
Members of the genus Cycas
ZipcodeZoo has pages for 221 species, subspecies, varieties, forms, and cultivars in this genus. Here are just 100 of them:
C. aculeata · C. acuminatissima · C. angulata (Marlborough Blue Sago) · C. annaikalensis (Annaikal Cycas (India)) · C. apoa · C. arenicola · C. armstrongii (Australian Cycad) · C. arnhemica · C. arnhemica arnhemica · C. arnhemica muninga · C. arnhemica natja · C. arnhemica subsp. muninga · C. arnhemica subsp. natja · C. aurea · C. avenicola · C. badensis · C. baguanheensis · C. balansae · C. basaltica · C. beddomei (Andhra Pradesh Cycas) · C. bellefonti · C. bifida (Fork-Leafed Cycad) · C. boddami · C. bougainvilleana (Cycad) · C. brachycantha · C. brevipinnata · C. brunnea · C. caffer · C. cairnsiana (Mount Surprise Sago) · C. calcicola · C. campestris (Cycad) · C. canalis · C. canalis canalis · C. canalis carinata · C. canalis subsp. carinata · C. candida · C. celebica · C. chamaoensis (Cycad) · C. chamberlainii · C. changjiangensis (Cycad) · C. chevalieri · C. chonarsu · C. circinalis (Palm-Leaved Cycas) · C. circinalis subsp. papuana · C. circinalis vera · C. circinnalis · C. clivicola (Mountain Cycad) · C. clivicola lutea · C. clivicola subsp. lutea · C. cochinchinensis · C. collina · C. comorensis · C. condaoensis · C. conferta · C. couttsiana (Cycad) · C. crassipes · C. cupida · C. curranii · C. debaoensis (Multipinnate Cycas) · C. desolata · C. diannanensis (Cycad) · C. dilatata · C. dolichophylla (Cycad) · C. edentata (Cycad) · C. edentata de · C. elephantipes (Cycad) · C. elongata · C. fairchildiana · C. fairylakea · C. falcata (Cycad) · C. ferruginea (Cycad) · C. fugax · C. furfuracea · C. glauca · C. glaucophylla · C. gracilis · C. gracilis var. glauca · C. gracilis var. viridis · C. guizhouensis · C. hainanensis (Hainan Cycas) · C. hamelini · C. hoabinhensis · C. hongheensis (Cycad) · C. hypoleuca · C. immersa · C. indica · C. inermis · C. intermedia · C. javana · C. jenkinsiana · C. kedia (Australian Nut Palm) · C. kennedyana · C. kirkii · C. lane-poolei · C. lindstromii · C. lingshuigensis · C. litoralis (Sea Sago) · C. longiconifera · C. longipetiolula · C. longisporophylla
More Info
- Search for Pictures: images.google.com
- Search for Scholarly Articles: Google Scholar
- Search using Scientific Name and Vernacular Names: All the Web | AltaVista Canada | AltaVista | Excite | Google | HotBot | Lycos
- Search using Specialized Databases: GenBank | Medline | Scirus | CISTI/CAL | Agricola Periodicals | Agricola Books
Further Reading
- Donaldson, J.S. (ed.) 2003. Cycads. Status Survey and Conservation Action Plan. IUCN/SSC Cycad Specialist Group. IUCN, Gland, Switzerland and Cambridge, UK.
- Fu Shu-hsia, Cheng Wan-chün, Fu Li-kuo & Chen Chia-jui. 1978. Cycadaceae. In: Cheng Wan-chün & Fu Li-kuo, eds., Fl. Reipubl. Popularis Sin. 7: 4-17.
Notes
Contributors
- African Regional Workshop (Conservation and Sustainable Management of Trees, Zimbabwe) 1998. Pericopsis elata. In: IUCN 2006. 2006 IUCN Red List of Threatened Species. <www.iucnredlist.org>. Downloaded on 20 October 2006.
- Barneby RC & Forero E, 1994/1995 (from ILDIS).
- Barneby RC & Grimes JW (Monograph 1996/97), 2001-05 (from ILDIS).
- Barneby RC and Forero E, 1994/1995 (from ILDIS).
- Bisby, F.A., Y.R. Roskov, M.A. Ruggiero, T.M. Orrell, L.E. Paglinawan, P.W. Brewer, N. Bailly, J. van Hertum, eds (2007). Species 2000 & ITIS Catalogue of Life: 2007 Annual Checklist. Species 2000: Reading, U.K.
- Brands, S.J. (comp.) 1989-2005. Systema Naturae 2000. The Taxonomicon. Universal Taxonomic Services, Amsterdam, The Netherlands. [http://sn2000.taxonomy.nl/Taxonomicon/]. Access date: Nov 23, 2005
- Brands, S.J. (comp.) 1989-2006. Systema Naturae 2000. The Taxonomicon. Universal Taxonomic Services, Amsterdam, The Netherlands. Accessed April 21, 2007.
- Brands, S.J. (comp.) 1989-2007. Systema Naturae 2000. The Taxonomicon. Universal Taxonomic Services, Amsterdam, The Netherlands. Accessed September 17, 2007.
- Carr, Gerald
- Chappill J, 2001-06 (from ILDIS).
- Cristofolini G, 2001-06 (from ILDIS).
- Fautin, Daphne G. (from Hexacorallians of the World).
- Global Biodiversity Information Facility. Accessed January 27, 2008. http://www.gbif.org Mediated distribution data from provider.
- Hexacorallians of the World 2001.
- Heyn CC & Heller D 1991/1993 (from ILDIS).
- Hill, K.D. 2003. In IUCN 2008. 2008 IUCN Red List of Threatened Species. IUCNRedList.org. Downloaded July 19, 2008.
- Home of the Xylariaceae 2006.
- ILDIS World Database of Legumes, 10, Nov 2005
- ILDIS World Database of LegumesNov 10, 2005.
- Lewis GP, 1994/1995 (from ILDIS).
- Light, Kris. East Tennessee Wildflowers
- Nghia, N.H. 1998. Dalbergia entadoides. In: IUCN 2006. 2006 IUCN Red List of Threatened Species. <www.iucnredlist.org>. Downloaded on 20 October 2006.
- Pippen, Jeffrey S. Jeff's Nature Page. Accessed December 8, 2007.
Data Sources
Accessed through GBIF Data Portal January 30, 2008:
- National Herbarium of New South Wales, NSW herbarium collection
Identifiers
- Biodiversity Heritage Library NamebankID: 3775998
- Catalogue of Life Accepted Name Code: IOP-200000-260
- Globally Unique Identifier: urn:lsid:ipni.org:names:297038-1
- International Plant Names Index (IPNI) ID: 297038-1
- IUCN ID: 42096
- Zipcode Zoo Species Identifier: 432755
Footnotes
- Jiarui Chen & Dennis W. Stevenson "Cycadaceae". in Flora of China Vol. 4 Page 1. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org. [back]
- "Cycas". in Flora of China Vol. 4 Page 1. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org. [back]
- Mean = -168.170 meters (-551.739 feet), Standard Deviation = 1,824.290 based on 6 observations. Altitude information for each observation from British Oceanographic Data Centre. [back]
