Overview
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Threatened |
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Common Names
Common Names in English:
Alpine Larkspur, Colorado Larkspur
Description
Family Ranunculaceae
Herbs perennial
or annual
, sometimes subshrubs
or herbaceous or woody vines
. Leaves basal and cauline, alternate, rarely opposite or whorled
, simple
or variously compound
, palmately nerved, rarely penninerved
, with or without stipules. Inflorescence a simple or compound monochasium, dichasium, simple or compound raceme, or flowers solitary. Flowers bisexual
, sometimes unisexual
, actinomorphic
, rarely zygomorphic, hypogynous. Sepals 3--6 or more, free
, petaloid
or sepaloid
, imbricate or sometimes valvate
in bud. Petals present or absent, 2--8 or more, free, usually with nectaries. Stamens numerous
, rarely few, free; filaments
linear
or filiform
; anthers
latrorse
, introrse
, or extrorse
; sometimes some sterile
stamens becoming staminodes. Carpels numerous or few, rarely 1, free, rarely connate
to various degrees
; ovary with 1 to many ovules. Fruit follicles or achenes, rarely capsules or berries
. Seeds small, with abundant endosperm and minute embryo.
About 60 genera and 2500 species: worldwide, but richly represented in N temperate regions
, particularly in E Asia; 38 genera (four endemic) and 921 species (604 endemic) in China.[1]
Genus Delphinium
Herbs, perennial
, from fasciculate roots
or rhizomes. Leaves basal and/or cauline, petiolate
, petioles
gradually to abruptly shorter on distal leaves; basal leaves
usually larger than cauline; cauline leaves alternate. Leaf blade
deeply palmately divided
, round to pentagonal
or reniform
, margins
entire or lobes
apically crenate
or lacerate
, lobes of basal blades wider and fewer than those of cauline blades. Inflorescences terminal
, 2-100(-more) -flowered racemes
(occasionally branched, thus technically panicles), 5-40 cm or more; bracts subtending
inflorescence branches; pedicels present or absent; bracteoles (on pedicels) subopposite-subalternate, not forming involucre. Flowers bisexual
, bilaterally symmetric
; sepals not persistent
in fruit, 5; upper sepal 1, spurred
, 8-24 mm; lateral
sepals 2, ± ovate
to elliptic
, 8-18 mm; lower sepals 2, similar to lateral sepals; upper petals 2, spurred, enclosed in upper sepal, nectary
inside tip
of spur; lower petals 2, plane
, ± ovate, ± 2-lobed, clawed, 2-12 mm, nectary absent; stamens 25-40; filaments
with base
expanded; staminodes absent between stamens and pistils; pistils 3(-5), simple
; ovules 8-20 per pistil; style present. Fruits follicles, aggregate, sessile, ± curved-cylindric, sides prominently veined or not; beak
terminal, straight, 2-4 mm.
Seeds dark brown to black (often appearing white because of air
in seed coat
cells
), rectangular to pyramidal
, often ± rough surfaced. x
= 8.
Species ca.
300: n temperate
and arctic
subtropical
and, in Eastern Hemisphere, tropical
mountains (s of equator in Africa) .
Three Eurasian species of Delphinium -- D . elatum Linnaeus, D . grandiflorum Linnaeus, and D . tatsienense Franchet--have been commonly cultivated in North America. Of the nonnative taxa, only D . elatum is sporadically naturalized
, as far as is known. Isolating mechanisms in Delphinium appear to be primarily ecological, geographic, and/or temporal
. Where these distinctions are disrupted, introgression often exists. Hybridization occurs regularly between certain taxa, particularly in areas of disturbance
(e.g.
, roadcuts, drainage
ditches, clearcuts) . The more common and easily recognized hybrids are included
in the key
.
Many names
have been misapplied in Delphinium . The few misapplied names
mentioned in discussions below refer to relatively widespread problems.[2]
Physical Description
Species Delphinium alpestre
Stems 5-25 cm; base
green, puberulent
. Leaves cauline, 5-20, on proximal
1/5 of stem at anthesis
; petiole
1-10 cm. Leaf blade
round to pentagonal
,
1.5-5 × 2-5 cm, puberulent; ultimate
lobes
3-15, width
2-11
mm.
Inflorescences 2-8-flowered; pedicel 1-4 cm, puberulent; bracteoles
1-3 mm from flowers, green, linear-lanceolate, 6-10 mm, puberulent.
Flowers: sepals dark blue, apex rounded
, puberulent, lateral
sepals
spreading
to forward pointing, 11-14 × 5-7 mm, spurs straight
except usually slightly down-curved at apex, varying from 20°
above to 20° below horizontal, 8-12 mm; lower petal blades ±
covering stamens, 4-6 mm, clefts
2-4 mm; hairs
sparse, mostly near
base of cleft, centered on inner lobes, white. Fruits 7-12 mm, 3.5-4
times longer
than wide, puberulent. Seeds unwinged; seed coat
cells
elongate
, surface roughened. [source]
Delphinium alpestre is very similar to D. ramosum, possibly divergent
from that taxon
only since the most recent glaciation of North America,
during which ancestors
of D. alpestre might have survived on peaks
above the ice, while ancestors of D. ramosum survived in valleys
below the ice. Since glaciation, D. ramosum apparently has migrated
upslope, near but not adjoining populations of D. alpestre. [source]
Habit: Forb/herb
Flowers: The flower has five petals which grow together to form a hollow flower with a spur at the end. Blooms in late spring to late summer. Attracts butterflies and hummingbirds. • Bloom Period: July. • Flower Color: dark blue
Foliage: Deeply lobed with 3-7 toothed , pointed lobes .
Size/Age/Growth
Size: under 6" tall.
Landscaping
Care: Deer resistant. Heat tolerant . Use slug bait as needed. Work bone meal around the plants . Treat mildew with a mildew spray or dusting sulfur. When flowers fade, cut spikes just above foliage , leaving a foot of old stem.
Habitat
Exposed talus
slopes
on high peaks; of conservation
concern; (3400-)3800
m
and above[3].
Biology
Reproduction
Duration: Perennial
Growth
Culture: Space 12-15" apart. Allow seedheads to dry on plant before collection . Propate from seed by directly sowing outdoors in fall or using a coldframe or unheated greenhouse in winter. Finish planting by mid-April.
Soil: Prefers cool, rich, moist soil with a summer mulch. Needs good winter drainage .
Sunlight: Sun Exposure: Full sun to partial shade.
Moisture: Water Requirements: Irrigate flowering plants freely at their base . Sprinkling may cause mildew .
Temperature: Cold Hardiness: 3a, 3b, 4a, 5a, 5b, 6a, 6b, 7a, 7b, 8a, 8b. (map)
Taxonomy
- Domain:
Eukaryota
(
)
- Whittaker & Margulis,1978
- eukaryotes
- Kingdom:
Plantae
(
)
- Haeckel, 1866
- Plants
- Subkingdom:
Viridaeplantae
(
)
- Cavalier-Smith, 1981
- Phylum:
Tracheophyta
(
)
- Sinnott, 1935 Ex Cavalier-Smith, 1998
- Vascular Plants
- Subphylum:
Euphyllophytina
(
)
- Infraphylum:
Radiatopses
(
)
- Kenrick & Crane, 1997
- Class:
Magnoliopsida
(
)
- Brongniart, 1843
- Dicotyledons
- Subclass:
Ranunculidae
(
)
- Takhtajan Ex Reveal, 1992
- Superorder:
Ranunculanae
(
)
- Takhtajan Ex Reveal, 1992
- Order:
Ranunculales
(
)
- Dumortier, 1829
- Family:
Ranunculaceae
(
)
- Adans., 1763, Nom. Cons.
- Buttercup Family
- Subfamily:
Trollioideae
(
)
- Tribe:
Delphinieae
(
)
- Genus:
Delphinium
(
)
- Linnaeus, Sp. Pl. 1: 530. 1753; Gen. Pl. ed 5, 236. 1754.
- Larkspur, delphinium [Greek delphinion, derived from delphin, possibly for fancied resemblance of flowers of some species to classical sculptures of dolphins]
- Specific epithet:
alpestre
- Rydberg, Bull. Torrey Bot. Club. 29: 146. 1902.
- Botanical name: - Delphinium alpestre
- Specific epithet:
alpestre
- Rydberg, Bull. Torrey Bot. Club. 29: 146. 1902.
- Genus:
Delphinium
(
- Tribe:
Delphinieae
(
- Subfamily:
Trollioideae
(
- Family:
Ranunculaceae
(
- Order:
Ranunculales
(
- Superorder:
Ranunculanae
(
- Subclass:
Ranunculidae
(
- Class:
Magnoliopsida
(
- Infraphylum:
Radiatopses
(
- Subphylum:
Euphyllophytina
(
- Phylum:
Tracheophyta
(
- Subkingdom:
Viridaeplantae
(
- Kingdom:
Plantae
(
Unambiguous Synonyms
- Delphinium ramosum Rydberg var. alpestre (Rydberg) W. A. Weber
- Delphinium ramosum var. alpestre (Rydb.) W. A. Weber
Notes
Name Status: Accepted Name . Latest taxonomic scrutiny: 15-Mar-2000
Similar Species
Members of the genus Delphinium
ZipcodeZoo has pages for 1488 species, subspecies, varieties, forms, and cultivars in this genus. Here are just 100 of them:
D. 'A.J. Moir' · D. 'Abendleuchten' · D. abietorum · D. aconiti · D. aconitifolium · D. aconitioides · D. actaea · D. acuminatissimum · D. acutilobum · D. addendum · D. aemulans · D. aff. crassifolium · D. afghanicum · D. 'After Midnight' · D. ageratum · D. 'Agnes Brookes' · D. 'Ailsa' · D. aitchisonii · D. ajacis · D. ajacis 'Lonnies' (Rocket Larkspur) · D. ajaus · D. aktoense · D. alabamicum (Alabama Larkspur) · D. alatum · D. albescens · D. albiflorum · D. albocoeruleum · D. albocoeruleum var. przewalskii · D. albomarginatum · D. 'Alice Artindale' · D. 'Alie Duyvensteyn' · D. 'Alison Claire' · D. alpestre (Colorado Larkspur) · D. 'Alphonse Karr' · D. alpinum · D. altaicum · D. altissimum · D. amabile · D. amabile amabile · D. amabile clarianum · D. amabile var. amabile · D. 'Amadeus' · D. amani · D. ambiguum · D. 'Ambrose Woods' · D. americanum · D. amoenum · D. 'Amos Perry' · D. 'Amour' · D. amplibracteatum · D. anatolicum · D. andersoni · D. andersonii (Anderson´s Larkspur) · D. andersonii var. andersonii (Anderson´s Larkspur) · D. andersoni andersoni · D. andersoni cognatum · D. andersoni var. andersoni · D. andersoni var. cognatum · D. andesicola (Chiricahua Mountains Larkspur) · D. andesicola amplum (Chiricahua Mountain Larkspur) · D. andesicola andesicola (Chiricahua Mountain Larkspur) · D. 'Angela Harbutt' · D. angustipaniculatum · D. angustirhombicum · D. 'Anne Kenrick' · D. 'Anne Page' · D. 'Ann Woodfield' · D. anomalum · D. 'Antares' · D. anthoroideum · D. anthriscifolium · D. antoninum (Anthony Peak Larkspur) · D. antonium · D. apetalum · D. 'Aphrodite' · D. apiculatum · D. 'Apollo' · D. araraticum · D. arcuatum · D. 'Ariel' · D. armeniacum · D. 'Astolat' (Delphinium) · D. Astolat Group · D. 'Atholl' · D. atropurpureum · D. attenuatum · D. aucheri · D. 'Audrey Mott' · D. 'Augenweide' · D. austriacum · D. autumnale · D. Avon strain · D. axilliflorum · D. azureum · D. 'Baby Doll' · D. bakeri (Baker´s Larkspur) · D. balansae · D. balcanicum · D. baoshanense · D. 'Barbara Nason'
More Info
- Search for Pictures: images.google.com
- Search for Scholarly Articles: Google Scholar
- Search using Scientific Name and Vernacular Names: All the Web | AltaVista Canada | AltaVista | Excite | Google | HotBot | Lycos
- Search using Specialized Databases: GenBank | Medline | Scirus | CISTI/CAL | Agricola Periodicals | Agricola Books
Further Reading
- Bulletin of the Torrey Botanical Club. 29 1902 New York: Torrey Botanical Club, 1870- ENG url p. 146.
- Contributions from the New York Botanical Garden. New York: The Garden, 1899- ENG url p. 101, p. 468.
- Flora of Colorado, by P.A. Rydberg. Fort Collins, Col., Experiment Station, 1906. ENG url p. 139, p. 139.
- Flora of New Mexico / by E.O. Wooton and Paul C. Standley. Washington: G.P.O., 1915. ENG url p. 244, p. 245.
- Just's botanischer jahresbericht. Systematisch geordnetes repertorium der botanischen literatur aller länder. Berlin, Gebr. Borntraeger, 1874-98; GER url p. 200.
- Ewan, J. 1945. A synopsis of the North American species of Delphinium. Univ. Colorado Stud., Ser. D, Phys. Sci. 2: 55-244.
- Lewis, H. and C. Epling. 1954. A taxonomic study of Californian delphiniums. Brittonia 8: 1-22.
- Taylor, R. J. 1960. The genus Delphinium in Wyoming. Univ. Wyoming Publ. 24: 9-21.
- Warnock, M. J. 1993. Delphinium. In: J. C. Hickman, ed. 1993. The Jepson Manual. Higher Plants of California. Berkeley, Los Angeles, and London. Pp. 916-922.
- Warnock, M. J. 1995. A taxonomic conspectus of North American Delphinium. Phytologia 78(2): 73-101.
Notes
Contributors
- Bisby, F.A., Y.R. Roskov, M.A. Ruggiero, T.M. Orrell, L.E. Paglinawan, P.W. Brewer, N. Bailly, J. van Hertum, eds (2007). Species 2000 & ITIS Catalogue of Life: 2007 Annual Checklist. Species 2000: Reading, U.K.
- Brands, S.J. (comp.) 1989-2006. Systema Naturae 2000. The Taxonomicon. Universal Taxonomic Services, Amsterdam, The Netherlands. Accessed April 19, 2007.
- "Delphinium alpestre". in Flora of North America Vol. 3. Published by Oxford University Press. Online at EFloras.org.
- Global Biodiversity Information Facility. Accessed February 28, 2008. http://www.gbif.org Mediated distribution data from 2 providers.
- USDA, NRCS. 2005. The PLANTS Database, Version 3.5 (http://plants.usda.gov). National Plant Data Center, Baton Rouge, LA 70874-4490 USA.
Data Sources
Accessed through GBIF Data Portal February 28, 2008:
- University of Colorado Museum: Zoological specimens
- USDA PLANTS: USDA PLANTS Database
Identifiers
- Biodiversity Heritage Library NamebankID: 2645375
- Catalogue of Life Accepted Name Code: ITS-18528
- Globally Unique Identifier: urn:lsid:ipni.org:names:77259-2
- Integrated Taxonomic Information System (ITIS) Taxonomic Serial Number (TSN): 18528
- Natural Heritage Network Species Identifier: PDRAN0B020
- U.S.D.A. Plant Symbol: DERAA
- Zipcode Zoo Species Identifier: 34621
Footnotes
- Wencai Wang, Dezhi Fu, Liang-Qian Li, Bruce Bartholomew, Anthony R. Brach, Bryan E. Dutton, Michael G. Gilbert, Yuichi Kadota, Orbélia R. Robinson, Michio Tamura, Michael J. Warnock, Guanghua Zhu & Svetlana N. Ziman "Ranunculaceae". in Flora of China Vol. 6 Page 133. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org. [back]
- Michael J. Warnock "Delphinium". in Flora of North America Vol. 3. Oxford University Press. Online at EFloras.org. [back]
- "Delphinium alpestre". in Flora of North America Vol. 3. Oxford University Press. Online at EFloras.org. [back]
