| Your Home Page | Contact Us |

• Conservation Status
• Taxonomy
• Physical Description
• Distribution
• Reproduction
• Similar Species
• Bibliography
• More Info
• Notes
• Keep Exploring

Overview:

Critically Endangered

Conservation Status

NatureServe: G1 Critically Imperiled

Taxonomy

Notes:

Name Status: Accepted Name. Latest taxonomic scrutiny: 15-Mar-2000

Physical Description

Family Boraginaceae:

Herbs perennial, biennial, or annual, less often lianas, shrubs, or trees, usually bristly or scabrous-pubescent. Leaves simple, exstipulate, alternate, rarely opposite, entire or serrate at margin. Inflorescences often double scorpioid cymes, rarely solitary; bracts present or absent. Flowers bisexual, actinomorphic, rarely zygomorphic. Calyx usually 5-parted or lobed, mostly persistent. Corolla tubular, campanulate, rotate, funnelform, or salverform; tube appendages 5, rarely more, mostly trapeziform, rarely absent, sometimes a ring of hairs present; limb usually 5-parted; lobes overlapping, rarely twisted in bud. Stamens 5, inserted on corolla tube or rarely at throat, included or rarely exserted; anthers introrse, 2-loculed, usually dorsifixed at base, less often medifixed, dehiscence longitudinal. Nectaries at base of corolla tube or on disc below ovary. Ovary superior, 2-carpellate; locules 2 and each with 2 ovules, or 4 and each with 1 ovule; ovules nearly atropous, semianatropous, or anatropous. Style terminal or gynobasic, branched or not. Gynobase flat, fastigiate, or subulate. Fruit 1-4-seeded drupes or nutlets (mericarps) ; nutlets mostly dry, often ornamented with wings, prickles and/or glochids (stiff bristles with barbed or anchorlike tips) . Seeds vertical or oblique, coat membranous; embryo straight, less often curved; cotyledons flat, fleshy.

About 156 genera and 2500 species: temperate and tropical regions, centered in the Mediterranean region; 47 genera and 294 species in China, of which four genera and 156 species are endemic.^[1]

Genus Hackelia:

Herbs perennial or annual, appressed strigose or villous, 20-100 cm tall. Leaves alternate, green, large, thin. Inflorescences rarely solitary, terminal, not branched or paniculate. Fruiting pedicel deflexed. Calyx 5-parted to base or nearly so; lobes enlarged in fruit, vertical to reflexed. Corolla blue, light blue, or light purple, rarely yellow or white, campanulate-rotate to campanulate-tubular; throat with appendages, rarely appendages obscure or absent; lobes 5, imbricate in bud, vertical or divaricate at anthesis. Stamens inserted on corolla, included; anthers orbicular, ovate to oblong. Style and stigma solitary, usually not exceeding nutlets. Gynobase short pyramidal, to 2-3 mm. Nutlets 4, all or some developed, turbinate to ovoid, triangular-ovoid and dorsally compressed, marginal rib with compressed triangular to lanceolate glochids.

About 45 species: mainly in temperate regions of the Northern Hemisphere, and Central and South America; three species in China.

Lian & Wang (Acta Phytotax. Sin. 18: 514-520. 1980; Fl. Reipubl. Popularis Sin. 64(2) : 116-151. 1989) argued that the differences between Eritrichium and Hackelia are insufficient to keep them as separate genera. However, the gynobase in Eritrichium is cushion-shaped, to 1 mm high, while in Hackelia it is distinctly conical and 2-3 mm high. It is likely that other species from China listed under Eritrichium will need to be transferred to Hackelia in the future.^[2]

Habit: Forb/herb

Images:

Distribution

Range and Population

North America

Reproduction

Duration: Perennial

Similar Species

Members of the genus Hackelia:

There are approximately 99 species, subspecies, varieties, forms, and cultivars in this genus: H. deflexa americana · H. amethystima · H. amethystina (Amethyst Stickseed) · H. andicola · H. angustifolia · H. arida var. cusickii · H. bella (Greater Showy Stickseed) · H. besseyi (Bessey Stickseed) · H. bhutanica · H. brachytuba · H. brevicula (Poison Canyon Stickseed) · H. caerulescens · H. californica (California Stickseed) · H. ciliata (Okanogan Stickseed) · H. cinerea (Gray Stickseed) · H. costaricensis · H. cottonii · H. cronquistii (Cronquist's Stickseed) · H. cusickii (Cusick Stickseed) · H. davisii (Davis Stickseed) · H. deflexa (Nodding Stickseed) · H. deflexa var. americana · H. dielsii · H. difformis · H. diffusa (Spreading Stickseed) · H. diffusa var. arida · H. diffusa var. cottonii · H. diffusa var. diffusa · H. eastwoodae · H. eastwoodiae · H. echinocarya · H. elegans · H. floribunda (Davis Mountain Stickseed) · H. floribunda var. pinetorum · H. glochidiata · H. gracilenta (Colorado Stickseed) · H. guatemalensis · H. heliocarpa · H. hendersoni · H. hendersoni var. trachyphylla · H. hendersonii · H. hendersonii var. trachyphylla · H. hirsuta (New Mexico Stickseed) · H. hispida (Rough Stickseed) · H. hispida var. disjuncta · H. hispida var. hispida · H. ibapensis (Deep Creek Stickseed) · H. leonotis · H. leptophylla var. besseyi · H. leucantha · H. macrophylla · H. matsudairai · H. meeboldii · H. mexicana · H. micrantha (Blue Stickseed) · H. minima · H. mundula (Pink Stickseed) · H. murgabica · H. nelsonii · H. nervosa (Sierra Stickseed) · H. nipponica · H. obtusifolia · H. ophioba · H. ophiobia (Owyhee River Stickseed) · H. pamirica · H. parviflora · H. patens (Common Stickseed) · H. patens var. harrisonii · H. patens var. patens · H. patens var. semiglabrz · H. pinetorum (Livermore Stickseed) · H. pinetorum var. jonesii · H. pinetorum var. pinetorum · H. popovii · H. rattanii · H. revoluta · H. revoluta f. fiebrigii · H. revoluta f. ovatifolia · H. revoluta var. ovatifolia · H. roylei · H. scaberrima · H. scaberrima var. angustata · H. setosa (Bristly Stickseed) · H. sharsmithii (Sharsmith's Stickseed) · H. skutchii · H. standleyi · H. stewartii · H. stricta · H. strigosa · H. thompsoni · H. thymifolia · H. uncinata · H. ursina (Chihuahuan Stickseed) · H. ursina var. diaboli · H. ursina var. pustulata · H. ursina var. ursina · H. velutina (Velvet Stickseed) · H. venusta (Lesser Showy Stickseed) · H. virginiana (Beggar's-Lice)

Bibliography

• Wang Wen-tsai, Liu Yu-lan, Zhu Ge-ling, Lian Yong-shan, Wang Jing-quan & Wang Qing-rui in Kung Xian-wu & Wang Wen-tsai, eds. 1989. Boraginaceae. Fl. Reipubl. Popularis Sin. 64(2): 1-253.

More Info

• Search for Pictures: images.google.com
• Search for Scholarly Articles: Google Scholar
• Search using Scientific Name and Vernacular Names: All the Web | AltaVista Canada | AltaVista | Excite | Google | HotBot | Lycos
• Search using Specialized Databases: GenBank | Medline | Scirus | CISTI/CA L | Agricola Periodicals | Agricola Books
• Translate this page using Google: Arabic | Chinese | Deutsch | Español | Français | Italiano | Japanese | Korean | Português

Notes

Contributors:

• Bisby, F.A., Y.R. Roskov, M.A. Ruggiero, T.M. Orrell, L.E. Paglinawan, P.W. Brewer, N. Bailly, J. van Hertum, eds (2007). Species 2000 & ITIS Catalogue of Life: 2007 Annual Checklist. Species 2000: Reading, U.K.
• Brands, S.J. (comp.) 1989-2005. Systema Naturae 2000. The Taxonomicon. Universal Taxonomic Services, Amsterdam, The Netherlands. [http://sn2000.taxonomy.nl/Taxonomicon/]. Access date: Nov 23, 2005
• Global Biodiversity Information Facility. Accessed February 28, 2008. http://www.gbif.org Mediated distribution data from 2 providers.
• USDA, NRCS. 2005. The PLANTS Database, Version 3.5 (http://plants.usda.gov). National Plant Data Center, Baton Rouge, LA 70874-4490 USA.

Data Sources:

Accessed through GBIF Data Portal February 28, 2008:

• USDA PLANTS: USDA PLANTS Database
• Utah Valley State College (UVSC): Utah Valley State College Herbarium

Identifiers:

• Integrated Taxonomic Information System (ITIS) Taxonomic Serial Number (TSN): 502864
• Global Biodiversity Information Facility Taxonkey: 290212
• Globally Unique Identifier: urn:lsid:ipni.org:names:897884-1
• Natural Heritage Network Species Identifier: PDBOR0G0V0
• U.S.D.A. Plant Symbol: HAIB
• Zipcode Zoo Species Identifier: 41655

Footnotes:

1. Gelin Zhu, Harald Riedl & Rudolf V. Kamelin "Boraginaceae". in Flora of China Vol. 16 Page 329. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org.
2. "Hackelia". in Flora of China Vol. 16 Page 390. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org.