Common Names
Common Names in English:
Mediterranean Barley, Sea Barley, Seaside Barley
Description
Family Poaceae
Annual
or perennial
herbs, or tall woody bamboos
. Flowering stems (culms
) jointed
, internodes hollow or solid; branches arising singly from nodes and subtended by a leaf sheath
and 2-keeled prophyll, often fascicled in bamboos. Leaves arranged alternately in 2 ranks
, differentiated into sheath, blade
, and an adaxial
erect
appendage
at sheath/blade junction (ligule) ; leaf sheath surrounding and supporting culm-internode, split to base
or infrequently tubular
with partially or completely fused margins
, modified with reduced blade in bamboos (culm sheaths) ; leaf blades divergent, usually long, narrow and flat, but varying from inrolled
and filiform
to ovate
, veins parallel, sometimes with cross-connecting veinlets
(especially in bamboos) ; ligule membranous or a line
of hairs
. Inflorescence terminal
or axillary
, an open, contracted
, or spikelike panicle, or composed of lax
to spikelike racemes
arranged along an elongate
central axis, or digitate, paired
, or occasionally solitary; axillary inflorescences often many, subtended by spatheoles
(specialized bladeless leaf sheaths) and gathered into a leafy compound
panicle; spikelets
often aggregated into complex
clusters
in bamboos. Spikelets composed of distichous bracts arranged along a slender axis (rachilla) ; typically 2 lowest bracts (glumes
) empty, subtending
1 to many florets
; glumes often poorly differentiated from accompanying bracts in bamboos. Florets composed of 2 opposing bracts enclosing a single small flower, outer bract (lemma) clasping
the more delicate, usually 2-keeled inner bract (palea) ; base of floret often with thickened prolongation articulated with rachilla (callus) ; lemma often with apical or dorsal bristle
(awn
), glumes also sometimes awned
. Flowers bisexual
or unisexual
; lodicules (small scales
representing perianth) 2, rarely 3 or absent, 3 to many in bamboos, hyaline
or fleshy
; stamens 3 rarely 1, 2, 6, or more in some bamboos, hypogynous, filaments
capillary
, anthers
versatile; ovary 1-celled, styles (1 or) 2(rarely 3), free
or united
at base, topped by feathery stigmas, exserted from sides or apex of floret. Fruit normally a dry indehiscent caryopsis with thin pericarp firmly adherent
to seed, pericarp rarely free, fleshy in some bamboos; embryo small or large; hilum
punctate
to linear
.
About 700 genera and 11,000 species: widely distributed in all regions of the world.[1]
Genus Hordeum
Plants
annual
or perennial
. Culms
usually erect
. Leaf sheath
of cauline leaves split almost to base
; ligule membranous or leathery-membranous; auricles
present or absent; leaf blade
usually flat. Spike dense, usually without a terminal
spikelet
; rachis short, brittle, rarely flexible
. Spikelets usually 3 per node, in regular rows
, with 1(or 2) florets
; lateral
spikelets usually pedicellate
, rarely sessile, often reduced and much smaller than central spikelet; central spikelet usually sessile, rarely pedicellate, perfect
. Glumes
narrow, subulate-setaceous, sometimes lanceolate dilated
at base, inconspicuously 1-3-veined, not keeled
. Lemma subrounded abaxially, leathery, rarely leathery-membranous, 5-veined, not keeled, awned
or awnless. Palea almost equaling lemma, glabrous
, scabrous
, or ciliate
along keels. Lodicules broadly lanceolate or lanceolate-ovate. Caryopsis usually adnate
to lemma and palea, rarely free
, oblong
, concave
furrowed
on inner side, apex hairy
. x = 7.
Between 30 and 40 species: temperate regions
, also on subtropical
mountains; ten species (one endemic, two introduced
) in China.
All species when young are used for forage
.[2]
Physical Description
Habit: Graminoid
Habitat
Typically found at an altitude of 0 to 4,557 meters (0 to 14,951 feet).[3]
Biology
Reproduction
Duration: Annual
Taxonomy
- Domain:
Eukaryota
(
)
- Whittaker & Margulis,1978
- eukaryotes
- Kingdom:
Plantae
(
)
- Haeckel, 1866
- Plants
- Subkingdom:
Viridaeplantae
(
)
- Cavalier-Smith, 1981
- Phylum:
Magnoliophyta
(
)
- Sinnott, 1935 Ex Cavalier-Smith, 1998
- Flowering Plants
- Subphylum:
Euphyllophytina
(
)
- Infraphylum:
Radiatopses
(
)
- Kenrick & Crane, 1997
- Class:
Liliopsida
(
)
- Scopoli, 1760
- Subclass:
Commelinidae
(
)
- Takhtajan, 1967
- Superorder:
Poanae
(
)
- (Small, 1903) Takhtajan, 1997 Ex Reveal & Doweld, 1999
- Superorder:
Poanae
(
- Subclass:
Commelinidae
(
- Class:
Liliopsida
(
- Infraphylum:
Radiatopses
(
- Subphylum:
Euphyllophytina
(
- Phylum:
Magnoliophyta
(
- Subkingdom:
Viridaeplantae
(
- Kingdom:
Plantae
(
Unambiguous Synonyms
- Critesion marinum /i> (Huds.) 脕. L枚ve
Notes
Name
Status: Accepted Name
. Latest taxonomic
scrutiny: 15-Mar-2000
Place of publication
: Fl.
angl. ed. 2, 1:57. 1778
Name verified on 05-Nov-2007 by ARS Systematic Botanists. Last updated: 05-Nov-2007
Similar Species
Members of the genus Hordeum
ZipcodeZoo has pages for 332 species, subspecies, varieties, forms, and cultivars in this genus. Here are just 100 of them:
H. adpressum · H. adscendens · H. aegiceras · H. aestivum · H. agriocrithon · H. ambiguum · H. andicola · H. andinum · H. anglicum · H. apertum · H. arenarium · H. arizonicum (Arizona Barley) · H. asperum · H. astrachanense · H. avenaceum · H. berteroanum · H. bifarium · H. blomii · H. bogdani · H. bogdanii · H. bonariense · H. boreale · H. brachyantherum (Meadow Barley) · H. brachyantherum brachyantherum (Northern Barley) · H. brachyantherum californicum (California Barley) · H. brachyantheum · H. brachyatherum · H. bracyantherum · H. breviaristatum · H. brevisubulatum · H. brevisubulatum var. hirtellum · H. brevisubulatum var. turkestanicum · H. bulbosum (Bulbous Barley) · H. bulbosum bulbosum (Bulbous Barley) · H. caducum · H. caespitosum · H. californica · H. californicum · H. canadense · H. capense · H. caput-medusae · H. cartilagineum · H. chilense · H. chilense var. magellanicum · H. chilense var. muticum · H. ciliatum · H. coeleste · H. coleophorum · H. comosum · H. comosum var. bifidum · H. comosum var. pubiflorum · H. complanatum · H. compressum · H. cordobense · H. crinitum · H. cylindricum · H. daghestanicum · H. deficiens · H. delileanum · H. delphicum · H. depauperatum · H. depilatum · H. depressum (Dwarf Barley) · H. dilatatum · H. distichon · H. distichon var. erectum 'Belfor' · H. distichum · H. divergens · H. durum · H. elisabethpolense · H. elongatum · H. elymoides · H. erectifolium · H. euclaston · H. euclaston var. pubescens · H. europaeum · H. flexicaule · H. flexuosum · H. fragile · H. fuegianum · H. fuscum · H. geniculatum · H. giganteum · H. glabrum · H. glaucum · H. guatemalense · H. gussoneanum · H. gussonianum · H. gymnodistichum · H. halophilum · H. halophilum var. breviaristatum · H. hexaploidum · H. hexastichon · H. hexastichum · H. himalayense · H. hirsutum · H. histrix · H. hohenackeri · H. horsfordianum · H. hrasdanicum
More Info
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Further Reading
- Berichte der Schweizerischen Botanischen Gesellschaft = Bulletin de la Socit botanique Suisse. [Zrich]: Die Gesellschaft, 1891-1981. FRE url p. 342.
- Botanisk tidsskrift / utgivet af den Botaniske forening i K鴅enhavn. K鴅enhavn: H. Hagerups Forlag, ENG url p. 466.
- Bothmer, R. von. 1996. Distribution and habitat preferences in the genus Hordeum in Iran and Turkey. Ann. Naturhist. Mus. Wien 98B(suppl.):113.
- Bulletin de l'Acadmie internationale de gographie botanique. Le Mans: Impr. Edmond Monnoyer, 1899-1910. FRE url p. 26.
- Caroli Ludovici Willdenow Florae Berolinensis prodromus: secundum systema Linneanum ab illustr. viro ac eq. C.P. Thunbergio emendatum conscriptus. Berolini: Impensis Wilhelmi Viewegii, 1787. LAT url p. 62.
- Correa, M. N. 1969–. Flora patagonica. (F Patag)
- Davis, P. H., ed. 1965–1988. Flora of Turkey and the east Aegean islands. (F Turk)
- Die Lepidopteren der Schweiz, von Prof. Dr. Heinrich Frey. Leipzig, W. Engelmann, 1880. GER url p. 45.
- FNA Editorial Committee. 1993–. Flora of North America. (F NAmer)
- Great Basin naturalist memoirs. [Provo, Utah]Brigham Young University, 1976-1992. ENG url p. 740, p. 741.
- Journal of botany, British and foreign. London: Robert Hardwicke, 1863-1942. ENG url p. 193, p. 260.
- Just's botanischer jahresbericht. Systematisch geordnetes repertorium der botanischen literatur aller l鋘der. Berlin, Gebr. Borntraeger, 1874-98; GER url p. 14, p. 29.
- Manual of the grasses of the United States / by A.S. Hitchcock. Washington: U.S. G.P.O., 1935. ENG url p. 266, p. 871.
- Nuova flora analitica d'Italia, contenente la descrizione delle piante vascolari indigene inselvatichite e largamente coltivate in Italia, del dott. Adriano Fiori. Firenze, Tipografia di M. Ricci, 1923-29. ITA url p. 162.
- R. von Bothmer & N. Jacobsen. 1985. Origin, taxonomy, and related species. In: American Society of Agronomy, Agronomy; a series of monographs. (Agron Monog) 26:47.
- R. von Bothmer et al. 1989. Variation and differentiation in Hordeum marinum (Poaceae). Nordic J. Bot. 9:6.
- Repertorium specierum novarum regni vegetabilis. Berlin: F. Fedde: 1910-1942. GER url p. 108.
- Skvortsov, A. K., ed. 2006–. Flora of the Lower Volga region. (F LowVolga)
- The Annals of Scottish natural history. Edinburgh: David Douglas, 1892-1911. ENG url p. 102, p. 112, p. 229, p. 34, p. 42.
- The Great Basin naturalist. Provo, Utah, M.L. Bean Life Science Museum, Brigham Young University. ENG url p. 38, p. 562, p. 95.
- Tutin, T. G. et al., eds. 1964–1980. Flora europaea. (F Eur)
- Tzvelev, N. N. 1976. Zlaki SSSR. (Grass USSR)
- Weber, E. 2003. Invasive plant species of the world: a reference guide to environmental weeds. (Invasive Pl Spec)
- Zuloaga, F. O. et al. 1994. Catalogo de la familia Poaceae en la Republica Argentina. Monogr. Syst. Bot. Missouri Bot. Gard. 47. (L Grass Argent)
- Chen Shouliang, Jin Yuexing, Zhuang Tide, Fang Wenzhe, Sheng Guoying, Liu Liang, Wu Zhenlan, Lu Shenglian, Sun Bisin, Hu Zhihao, Wang Song, Sun Xiangzhong, Wang Huiqin, Yang Xilin, Wang Chaopin, Li Binggui & Wen Shaobin. 1990. Gramineae (Poaceae) (4). In: Chen Shouliang, ed., Fl. Reipubl. Popularis Sin. 10(1):1401
- Chen Shouliang, Zhuang Tide, Fang Wenzhe, Sheng Guoying, Jin Yuexing, Liu Liang, Sun Bisin, Hu Zhihao & Wang Song. 1997. Gramineae (Poaceae) (5). In: Chen Shouliang, ed., Fl. Reipubl. Popularis Sin. 10(2): 1301
- Liu Liang, Zhu Taiping, Chen Wenli, Wu Zhenlan & Lu Shenglian. Gramineae (Poaceae) (2). In: Liu Liang, ed., Fl. Reipubl. Popularis Sin. 9(2): 1405
- Lu Sheng-lian, Sun Yong-hua, Liu Shang-wu, Yang Yong-chang, Wu Zhen-lan, Kuo Pen-chao, Yang Hsi-ling, Wang Chao-pin & Tsui Nai-ran. 1987. Gramineae (3). In: Kuo Pen-chao, ed., Fl. Reipubl. Popularis Sin. 9(3): 1329
- Wang Zhengping, Ye Guanghan, Yang Yaling, Yu Zehua, Hu Chenhua, Geng Bojie, Feng Xuelin, Jia Liangzhi, Xia Nianhe, Li Dezhu, Zhang Weiping, Xue Jiru, Zhu Zhengde, Zhao Qiseng, Chen Shouliang, Sheng Guoying, Chen Shaoyun, Yao Changyu, Lu Jionglin, Sun Jiliang, Lin Wantao, Yi Tongpei, Zhao Huiru, Wen Taihui & Dai Qihui. 1996. Gramineae (Poaceae) (1).
Notes
Contributors
- Bisby, F.A., Y.R. Roskov, M.A. Ruggiero, T.M. Orrell, L.E. Paglinawan, P.W. Brewer, N. Bailly, J. van Hertum, eds (2007). Species 2000 & ITIS Catalogue of Life: 2007 Annual Checklist. Species 2000: Reading, U.K.
- Global Biodiversity Information Facility. Accessed March 29, 2007. http://www.gbif.org Mediated distribution data from provider.
- USDA, ARS, National Genetic Resources Program. Germplasm Resources Information Network - (GRIN) [Online Database]. National Germplasm Resources Laboratory, Beltsville, Maryland. URL (April 28, 2008)
Data Sources
Accessed through GBIF Data Portal November 29, 2007:
- Australian National Herbarium
- , Australian National Herbarium
- Bundesamt f眉r Naturschutz / Zentralstelle f眉r Phytodiversit盲t Deutschland, Bundesamt fuer Naturschutz / Zentralstelle fuer Phytodiversitaet Deutschland
- Comisi贸n nacional para el conocimiento y uso de la biodiversidad, Herbario del Instituto de Ecolog铆a, A.C., M茅xico
- Conservatoire botanique national du Bassin parisien, Conservatoire botanique national du Bassin parisien
- European Environment Agency, EUNIS
- GBIF-Spain, Bot谩nica, Universidad de Le贸n: LEB-Cormo
- GBIF-Spain, Direcci贸n General de Investigaci贸n, Desarrollo Tecnol贸gico e Innovaci贸n de la Junta de Extremadura(DGIDTI): HSS
- GBIF-Spain, Herbario Universidad de M谩laga: MGC-Corm贸fitos
- GBIF-Spain, Hortus Botanicus Sollerensis Herbarium
- GBIF-Spain, Jard铆n Bot谩nico de C贸rdoba: Herbarium COA
- GBIF-Spain, Real Jardin Botanico
- , Vascular Plant Herbarium
- GBIF-Spain, Universidad de Almer铆a, HUAL
- GBIF-Spain, Universidad de Extremadura, UNEX
- GBIF-Spain, Universidad de Oviedo. Departamento de Biolog铆a de Organismos y Sistemas: FCO
- International Plant Genetic Resources Institute(IPGRI), EURISCO
- International Plant Genetic Resources Institute(IPGRI), The System-wide Information Network for Genetic Resources
- Israel Nature and Parks Authority, Israel Nature and Parks Authority
- Leibniz Institute of Plant Genetics and Crop Plant Research
- , IPK Genebank
- Missouri Botanical Garden, Missouri Botanical Garden
- NLBIF, Limnodata
- The Swedish Museum of Natural History
- , Lund Botanical Museum
- UK National Biodiversity Network, Botanical Society of the British Isles - Vascular Plants Database
- UK National Biodiversity Network, Joint Nature Conservation Committee - Vegetation surveys of coastal shingle in Great Britain
- USDA PLANTS, USDA PLANTS Database
- University of Vienna, Institute for Botany - Herbarium WU, Herbarium WU
- Utah State University, USU-UTC Specimen Database
- Utah Valley State College
- , Utah Valley State College Herbarium
Identifiers
- Biodiversity Heritage Library NamebankID: 2661066
- Catalogue of Life Accepted Name Code: ITS-40880
- Global Biodiversity Information Facility Taxonkey: 13793661
- Globally Unique Identifier: urn:lsid:ipni.org:names:76922-3
- GRIN Nomen Number: 19312
- Integrated Taxonomic Information System (ITIS) Taxonomic Serial Number (TSN): 40880
- Natural Heritage Network Species Identifier: PMPOA38090
- U.S.D.A. Plant Symbol: HOMAM
- Zipcode Zoo Species Identifier: 43108
Footnotes
- Shou-liang Chen, De-Zhu Li, Guanghua Zhu, Zhenlan Wu, Sheng-lian Lu, Liang Liu, Zheng-ping Wang, Bi-xing Sun, Zheng-de Zhu, Nianhe Xia, Liang-zhi Jia, Zhenhua Guo, Wenli Chen, Xiang Chen, Yang Guangyao, Sylvia M. Phillips, Chris Stapleton, Robert J. Soreng, Susan G. Aiken, Nikolai N. Tzvelev, Paul M. Peterson, Stephen A. Renvoize, Marina V. Olonova & Klaus Ammann "Poaceae". in Flora of China Vol. 22. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org. [back]
- "Hordeum". in Flora of China Vol. 22 Page 6, 386, 395, 443, 444. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org. [back]
- Mean = 212.220 meters (696.260 feet), Standard Deviation = 428.880 based on 1,164 observations. Altitude information for each observation from British Oceanographic Data Centre. [back]
