Overview
Ornamental shrub from tropical Africa with leaves 3-4 compound , often coppery or purplish; flowers purplish to red; and fruit dark red, about half an inch in diameter.
Common Names
Common Names in Portuguese (Brazil):
Léia, Léia-Alaranjada
Description
Family Leeaceae
Shrubs
, small trees
, scramblers
, or rarely large perennial
herbs. Stems unarmed
or with rows
of prickles; tendrils
absent. Leaves 1-4-pinnate to 3-foliolate or simple
; stipules sheathing
petiole
margins
with conspicuous
, persistent
or caducous
stipular
wings; leaflets
glabrous
to pubescent
with simple hairs, crenate
to serrate to dentate
at margin, teeth with small glandular
apex, lower surface usually with specialized multicellular
, stellate
or globular caducous "pearl" glands
. Inflorescences paniculate
, often corymbiform
, terminal
or axillary
, erect
or pendulous. Flowers hermaphroditic
, (4 or) 5-merous. Calyx campanulate
with triangular lobes
and glandular tips. Petals valvate
, apically often cucullate
, reflexed
at anthesis
, basally connate
, adnate
to staminodial tissue
and lower portion of floral
disk; floral disk tubular
, intrastaminal
. Stamens 4 or 5, antepetalous
, alternating with lobes of floral disk; anthers
tetrasporangiate
and 2-locular, introrse
and sometimes appearing extrorse
. Ovary superior but sometimes partly sunken
in disk, 2-3(-5) -carpellate but with a secondary septum in each carpel and 4-6(-10) -locular; ovule 1 per locule, anatropous
, bitegmic and crassinucellate
; style elongate
; stigma discoid
and capitate. Fruit a berry, rather dry, depressed
subglobose, purple, black, or orange. Seeds endotestal; endosperm ruminate
with roughly 5 ingrowths; embryo linear
. 2n = 24 or 48 (rarely 20, 22, or 28) .
One genus and ca.
34 species; widespread in tropical
and subtropical
Asia, extending into the Himalayan region and Australia, two species in Africa and Madagascar; ten species (two endemic) in China.[1]
Genus Leea
Erect
shrubs
or small trees
. Leaves alternate, 1-4-pinnate, rarely simple
or 3-foliolate. Inflorescences compound
dichasial or umbelliform. Flowers 4- or 5-merous, bisexual
. Lower part of petals adnate
to staminodial tube
on disk. Apex of staminodial tube 5-lobed, connate
to each other by thinner tissue
to form sinuses; apex of lobes
retuse
or 2-lobed, retusely apiculate
to bifid. Filaments
flattened, light brown, alternating with staminodial lobes, curved
inward; anthers
inverted
and packed within staminodial tube in bud; filaments straight and anthers long-exserted in open flower. Disk deeply cupulate
. Ovary discoid
, 4-6(-10) -locular; ovule 1 per locule; style short; stigma slightly thickened. Berry depressed-subglobose, 4-6(-10) -seeded. Endosperm ruminate
.
Geographic distribution is the same as that of the family
.[2]
Physical Description
Habit: Evergreen .
Size/Age/Growth
Size: 6-8' tall.
Biology
Growth
Sunlight: Sun Exposure: Light Shade.
Temperature: Cold Hardiness: 9a, 9b, 10a, 10b. (map)
Taxonomy
- Domain:
Eukaryota
(
)
- Whittaker & Margulis,1978
- eukaryotes
- Kingdom:
Plantae
(
)
- Haeckel, 1866
- Plants
- Subkingdom:
Viridaeplantae
(
)
- Cavalier-Smith, 1981
- Phylum:
Tracheophyta
(
)
- Sinnott, 1935 Ex Cavalier-Smith, 1998
- Vascular Plants
- Subphylum:
Euphyllophytina
(
)
- Infraphylum:
Radiatopses
(
)
- Kenrick & Crane, 1997
- Class:
Magnoliopsida
(
)
- Brongniart, 1843
- Dicotyledons
- Class:
Magnoliopsida
(
- Infraphylum:
Radiatopses
(
- Subphylum:
Euphyllophytina
(
- Phylum:
Tracheophyta
(
- Subkingdom:
Viridaeplantae
(
- Kingdom:
Plantae
(
Notes
An accepted name
in the RHS Horticultural Database.
Place of publication
: Gen. hist. 1:712. 1831
Name
verified on 22-Aug-2006 by ARS Systematic Botanists. Last updated: 22-Aug-2006
Similar Species
Members of the genus Leea
ZipcodeZoo has pages for 126 species, subspecies, varieties, forms, and cultivars in this genus. Here are just 100 of them:
L. aculeata · L. acuminata · L. acuminatissima · L. adwivedica · L. aequata · L. agusanensis · L. alata · L. amabilis · L. angulata · L. arborea · L. asiatica · L. aspera · L. banahaensis · L. biserrata · L. bracteata · L. brunoniana · L. bulusanensis · L. capitata · L. catanduanensis · L. cinerea · L. coccinea (West Indian Holly) · L. coccinea 'Rubra' (West Indian Holly) · L. compactiflora · L. congesta · L. cordata · L. coryphantha · L. crispa · L. cumingii · L. curtisii · L. cuspidifera · L. dentata · L. dielsii · L. diffusa · L. divaricata · L. edgeworthii · L. erecta · L. euphlebia · L. expansa · L. forbesii · L. fuliginosa · L. gigantea · L. glabra · L. gonioptera · L. gracilis · L. grandifolia · L. guineense · L. guineensis · L. guineensis 'Burgundy' · L. herbacea · L. heterodoxa · L. hirsuta · L. hirta · L. hispida · L. horrida · L. indica · L. integrifolia · L. javanica · L. krukoffiana · L. latifolia · L. linearifolia · L. longifolia · L. longifoliola · L. longipetiolata · L. lucida · L. luzonensis · L. macrophylla · L. macropus · L. magnifolia · L. manillensis · L. micholitzii · L. monophylla · L. naumanni · L. negrosense · L. negrosensis · L. nitida · L. nova-guineensis · L. ottilis · L. palawanensis · L. palembanica · L. pallida · L. pallidifolia · L. papillosa · L. papuana · L. parallela · L. parva · L. parvifoliola · L. pauciflora · L. philippinensis · L. pinnata · L. platyphylla · L. polyphylla · L. pumila · L. pycnantha · L. quadrifida · L. ramosii · L. robusta · L. rodatzii · L. roehrsiana · L. rubra (West Indian Holly) · L. sambucina
More Info
- Search for Pictures: images.google.com
- Search for Scholarly Articles: Google Scholar
- Search using Scientific Name and Vernacular Names: All the Web | AltaVista Canada | AltaVista | Excite | Google | HotBot | Lycos
- Search using Specialized Databases: GenBank | Medline | Scirus | CISTI/CAL | Agricola Periodicals | Agricola Books
Further Reading
- Annuaire du Conservatoire et du jardin botaniques de Genve. Genve: Georg & Cie, 1897-1919/1922. FRE url p. 91.
- Bulletin of the British Museum (Natural History). London: The Museum, 1951-1992. ENG url p. 365.
- Carrisso, L. W. et al., eds. 1937–. Conspectus florae angolensis. (F Angol)
- Exell, A. W. et al., eds. 1960–. Flora zambesiaca. (F Zamb)
- Grierson, A. J. C. & D. J. Long. 1984–. Flora of Bhutan including a record of plants from Sikkim. (F Bhutan)
- Hajra, P. K. et al., eds. 1999–. Flora of Andaman-Nicobar Islands. (F Andaman Nicobar)
- Huxley, A., ed. 1992. The new Royal Horticultural Society dictionary of gardening. (Dict Gard)
- Judd, W.S., Campbell, C.S., Kellog, E.A. and Donoghue, M.J. (2002): Plant Systematics: a phylogenetic approach, Sinauer, Sunderland, Mass.
- Just's botanischer jahresbericht. Systematisch geordnetes repertorium der botanischen literatur aller länder. Berlin, Gebr. Borntraeger, 1874-98; GER url p. 243, p. 597.
- Keay, R. W. J. & F. N. Hepper. 1953–1972. Flora of west tropical Africa, ed. 2. (F WT Afr)
- Liberia, by Sir Harry Johnston. With an appendix on the flora of Liberia, by Dr. Otto Stapf; 28 coloured illustrations by Sir Harry Johnston, 24 botanical drawings by Miss Matilda Smith, 402 black and white il London, Hutchinson, 1906. ENG url p. 590.
- Ridsdale, C. E. 1974. A revision of the family Leeaceae. Blumea 22:93.
- Sharma, B. D. et al., eds. 1993–. Flora of India. (F India)
- Staples, G. W. & D. R. Herbst. 1996. Nomenclatural changes affecting cultivated plants. I. Baileya 23:176.
- Steenis, C. G. G. J. van, ed. 1948–. Flora malesiana. (F Males)
- Turrill, W. B. et al., eds. 1952–. Flora of tropical East Africa. (F TE Afr)
- Li Chaoluan. 1998. Leea. In: Li Chaoluan, ed., Fl. Reipubl. Popularis Sin. 48(2): 3-12.
- Li Chaoluan. 1998. Leea. In: Li Chaoluan, ed., Fl. Reipubl. Popularis Sin. 48(2): 3-12.
Notes
Contributors
- Global Biodiversity Information Facility. Accessed December 01, 2007. http://www.gbif.org Mediated distribution data from 5 providers.
- USDA, ARS, National Genetic Resources Program. Germplasm Resources Information Network - (GRIN) [Online Database]. National Germplasm Resources Laboratory, Beltsville, Maryland. URL (May 04, 2008)
Data Sources
Accessed through GBIF Data Portal December 01, 2007:
- Herbarium of the University of Aarhus, The AAU Herbarium Database
- Missouri Botanical Garden, Missouri Botanical Garden
- Taiwan Biodiversity Information Facility, Magnoliophyta
Identifiers
- Biodiversity Heritage Library NamebankID: 3477736
- Global Biodiversity Information Facility Taxonkey: 16077911
- GRIN Nomen Number: 315504
- International Plant Names Index (IPNI) ID: 68307-1
- Zipcode Zoo Species Identifier: 707253
Footnotes
- Prof. Zhiduan Chen & Jun Wen "Leeaceae". in Flora of China Vol. 12 Page 115, 169. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org. [back]
- "Leea". in Flora of China Vol. 12 Page 169, 173. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org. [back]
