Overview
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Vulnerable |
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Interesting Facts
Common Names
Common Names in English:
American Bugleweed, American Water Horehound, American Waterhorehound, Bugleweed, Carolina Bugleweed, Carolina Water Horehound, Clasping Water Horehound, Common Water Horehound, Cut-Leaf Water-Horehound, European Bugleweed, Green Gromwell, Green Stoneseed, Gypsywort, Northern Bugleweed, Northern Water-Horehound, Oneflower Bugleweed, Rough Bugleweed, Rough Water-Horehound, Sessile-Leaved Bugleweed, Sherard's Waterhorehound, Southern Bog Bugleweed, Taperleaf Bugleweed, Taperleaf Water Horehound, Water Horehound, Waterhorehound
Description
Family Boraginaceae
Herbs perennial
, biennial, or annual
, less often lianas, shrubs
, or trees
, usually bristly
or scabrous-pubescent. Leaves simple
, exstipulate
, alternate, rarely opposite, entire or serrate at margin
. Inflorescences often double
scorpioid cymes, rarely solitary; bracts present or absent. Flowers bisexual
, actinomorphic
, rarely zygomorphic. Calyx usually 5-parted or lobed
, mostly persistent
. Corolla tubular
, campanulate
, rotate, funnelform
, or salverform
; tube
appendages
5, rarely more, mostly trapeziform, rarely absent, sometimes a ring
of hairs
present; limb usually 5-parted; lobes
overlapping, rarely twisted in bud. Stamens 5, inserted
on corolla tube or rarely at throat
, included
or rarely exserted; anthers
introrse
, 2-loculed, usually dorsifixed
at base
, less often medifixed
, dehiscence longitudinal
. Nectaries at base of corolla tube or on disc below ovary. Ovary superior, 2-carpellate; locules 2 and each with 2 ovules, or 4 and each with 1 ovule; ovules nearly atropous
, semianatropous, or anatropous
. Style terminal
or gynobasic
, branched or not. Gynobase
flat, fastigiate
, or subulate
. Fruit 1-4-seeded drupes or nutlets
(mericarps) ; nutlets mostly dry, often ornamented with wings
, prickles and/or glochids (stiff bristles
with barbed
or anchorlike tips
) . Seeds vertical
or oblique
, coat
membranous; embryo straight, less often curved
; cotyledons flat, fleshy
.
About 156 genera and 2500 species: temperate
and tropical regions
, centered in the Mediterranean region; 47 genera and 294 species in China, of which four genera and 156 species are endemic.[1]
Genus Lithospermum
Herbs annual
or perennial
, short strigose
. Leaves alternate. Cymes terminal
or flowers solitary, bracteate
. Calyx 5-parted nearly to base
, slightly enlarged in fruit. Corolla white, yellow, or violet, actinomorphic
, funnelform
or salverform
; throat
with appendages
or bands
of hairs
, or longitudinally crispate
; limb usually campanulate
, 5-parted; lobes
spreading
. Stamens included
; filament
very short; anthers
oblong-linear, apex obtuse
, mucronulate
. Style filiform
, not exserted; stigma entire or indistinctly 2-cleft, capitate. Gynobase
flat. Nutlets
white or gray, ovoid
, smooth
, shiny or tuberculate
; attachment scar
at base adaxially.
About 50 species: Africa, Asia, Europe, North America, South America; five species in China.[2]
Physical Description
Habit: Subshrub , Shrub , Forb/herb
Flowers: Bloom Period: June. • Flower Color: near white, white
Size/Age/Growth
Size: 18-24" tall.
Habitat
Typically found at an altitude of 0 to 2,998 meters (0 to 9,836 feet).[3]
Biology
Reproduction
Duration: Perennial
Growth
Culture: Space 6-9" apart.
Soil: Minimum pH: 6.1 • Maximum pH: 7.8
Sunlight: Sun Exposure: Full Sun .
Temperature: Cold Hardiness: 3a, 3b, 4a, 4b, 5a, 5b, 6a, 6b, 7a, 7b, 8a, 8b, 9a, 9b, 10a, 10b, 11. (map)
Taxonomy
- Domain:
Eukaryota
(
)
- Whittaker & Margulis,1978
- eukaryotes
- Kingdom:
Plantae
(
)
- Haeckel, 1866
- Plants
- Subkingdom:
Viridaeplantae
(
)
- Cavalier-Smith, 1981
- Phylum:
Tracheophyta
(
)
- Sinnott, 1935 ex Cavalier-Smith, 1998
- Vascular Plants
- Subphylum:
Euphyllophytina
(
)
- Infraphylum:
Radiatopses
(
)
- Kenrick & Crane, 1997
- Class:
Spermatopsida
(
)
- Brongniart, 1843
- Subclass:
Asteridae
(
)
- Takhtajan, 1967
- Superorder:
Solananae
(
)
- R. Dahlgren Ex Reveal, 1992
- Family:
Boraginaceae
(
)
- A.L. de Jussieu, 1789, nom. cons.
- borage, bourraches
- Subfamily:
Boraginoideae
(
)
- Tribe:
Lithospermeae
(
)
- Genus:
Lithospermum
(
)
- C. Linnaeus, 1753
- Stoneseed
- Specific epithet:
viride
- Greene
- Botanical name: - Lithospermum viride Greene
- Specific epithet:
viride
- Greene
- Genus:
Lithospermum
(
- Tribe:
Lithospermeae
(
- Subfamily:
Boraginoideae
(
- Family:
Boraginaceae
(
- Superorder:
Solananae
(
- Subclass:
Asteridae
(
- Class:
Spermatopsida
(
- Infraphylum:
Radiatopses
(
- Subphylum:
Euphyllophytina
(
- Phylum:
Tracheophyta
(
- Subkingdom:
Viridaeplantae
(
- Kingdom:
Plantae
(
Notes
Name
Status: Accepted Name
.
Last scrutiny: 15-Mar-2000
Similar Species
Members of the genus Lithospermum
ZipcodeZoo has pages for 27 species, subspecies, varieties, forms, and cultivars in this genus:
L. arvense arvense (Field Gromwell) · L. californicum (California Gromwell) · L. calycosum (Chinati Gromwell) · L. canescens (Gromwell) · L. caroliniense (Carolina Gromwell) · L. caroliniense var. caroliniense (Carolina Puccoon) · L. caroliniense var. croceum (Carolina Puccoon) · L. cobrense (Mountain Gromwell) · L. confine (Arizona Gromwell) · L. diffusum (Scrambling Gromwell) · L. erythrorhizon (Lithospermum) · L. incisum (Fringed Gromwell) · L. latifolium (American Gromwell) · L. matamorense (Rough Gromwell) · L. mirabile (San Antonio Gromwell) · L. multiflorum (Many-Flowered Gromwell) · L. obtusifolium (Rounded-Leaf Gromwell) · L. officinale (European Gromwell) · L. parksii (Parks Stoneseed) · L. parksii I.M.Johnst. var. parksii I.M.Johnst. (Parks' Gromwell) · L. parksii I.M.Johnst. var. rugulosum I.M.Johnst. (Parks' Gromwell) · L. parksii var. parksii (Parks' Stoneseed) · L. parksii var. rugulosum (Parks Stoneseed) · L. purpureocoeruleum (Gromwell) · L. ruderale (Columbia Puccoon) · L. tuberosum (Tuberous Gromwell) · L. viride (American Bugleweed)
More Info
- Search for Pictures: images.google.com
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Further Reading
- Contributions from the United States National Herbarium 13 1909-1912 Washington, D.C.: Smithsonian Institution Press, 1890- url p. 157, p. 211, p. 543.
- Flora of New Mexico / by E.O. Wooton and Paul C. Standley. Washington: G.P.O., 1915. url p. 543.
- Leaflets of western botany. San Fransisco:[J. T. Howell], 1932-1966. url .
- Synoptical flora of North America: the Gamopetalæ, being a second edition of vol. I, part II, and vol. II, part I, collected. London, Iverson, Blakeman, Taylor, and Company;1886. url .
- The Great Basin naturalist. 39 1979 Provo, Utah: M.L. Bean Life Science Museum, Brigham Young University, 1939-1999. url p. 307.
- Wang Wen-tsai, Liu Yu-lan, Zhu Ge-ling, Lian Yong-shan, Wang Jing-quan & Wang Qing-rui in Kung Xian-wu & Wang Wen-tsai, eds. 1989. Boraginaceae. Fl. Reipubl. Popularis Sin. 64(2): 1-253.
Notes
Contributors
Data Sources
Accessed through GBIF Data Portal November 20, 2007:
- Comisión nacional para el conocimiento y uso de la biodiversidad, Herbario del Instituto de Ecología, A.C., México
- School of Life Sciences, Arizona State University, Arizona State University Vascular Plant Herbarium
- USDA PLANTS, USDA PLANTS Database
Identifiers
- Biodiversity Heritage Library NamebankID: 2654754
- Catalogue of Life Accepted Name Code: ITS-31943
- Global Biodiversity Information Facility Taxonkey: 13749731
- Globally Unique Identifier: urn:lsid:ipni.org:names:118259-1
- Integrated Taxonomic Information System (ITIS) Taxonomic Serial Number (TSN): 32257
- Natural Heritage Network Species Identifier: PDLAM0X080 PDLAM0X030 PDLAM0X0A0 PDLAM0X060 PDLAM0
- U.S.D.A. Plant Symbol: LYVIP LYSH
- Zipcode Zoo Species Identifier: 47536
Footnotes
- Gelin Zhu, Harald Riedl & Rudolf V. Kamelin "Boraginaceae". in Flora of China Vol. 16 Page 329. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org. [back]
- "Lithospermum". in Flora of China Vol. 16 Page 342. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org. [back]
- Mean = 152.230 meters (499.442 feet), Standard Deviation = 235.890 based on 12,559 observations. Altitude information for each observation from British Oceanographic Data Centre. [back]
