Interesting Facts
- This is a noxious arable weed , widespread and naturalized in warm-temperate parts of the world (Darnel). The grains are often infected by a fungus (ergot) which produces the alkaloid temulin, causing poisoning when grazed by cattle or when present as a contaminant of flour . [source]
Common Names
Click on the language to view common names.
Common Names in English:
Bearded Darnel, Bearded Ryegrass, Darnel, Darnel Lolium Temulentum, Darnel Ryegrass, Poison Darnel
Common Names in French:
Ivraie énivrante
Common Names in German:
Taumellolch
Description
Family Poaceae
Annual
or perennial
herbs, or tall woody bamboos
. Flowering stems (culms
) jointed
, internodes hollow or solid; branches arising singly from nodes and subtended by a leaf sheath
and 2-keeled prophyll, often fascicled in bamboos. Leaves arranged alternately in 2 ranks
, differentiated into sheath, blade
, and an adaxial
erect
appendage
at sheath/blade junction (ligule) ; leaf sheath surrounding and supporting culm-internode, split to base
or infrequently tubular
with partially or completely fused margins
, modified with reduced blade in bamboos (culm sheaths) ; leaf blades divergent, usually long, narrow and flat, but varying from inrolled
and filiform
to ovate
, veins parallel, sometimes with cross-connecting veinlets
(especially in bamboos) ; ligule membranous or a line
of hairs
. Inflorescence terminal
or axillary
, an open, contracted
, or spikelike panicle, or composed of lax
to spikelike racemes
arranged along an elongate
central axis, or digitate, paired
, or occasionally solitary; axillary inflorescences often many, subtended by spatheoles
(specialized bladeless leaf sheaths) and gathered into a leafy compound
panicle; spikelets
often aggregated into complex
clusters
in bamboos. Spikelets composed of distichous bracts arranged along a slender axis (rachilla) ; typically 2 lowest bracts (glumes
) empty, subtending
1 to many florets
; glumes often poorly differentiated from accompanying bracts in bamboos. Florets composed of 2 opposing bracts enclosing a single small flower, outer bract (lemma) clasping
the more delicate, usually 2-keeled inner bract (palea) ; base of floret often with thickened prolongation articulated with rachilla (callus) ; lemma often with apical or dorsal bristle
(awn
), glumes also sometimes awned
. Flowers bisexual
or unisexual
; lodicules (small scales
representing perianth) 2, rarely 3 or absent, 3 to many in bamboos, hyaline
or fleshy
; stamens 3 rarely 1, 2, 6, or more in some bamboos, hypogynous, filaments
capillary
, anthers
versatile; ovary 1-celled, styles (1 or) 2(rarely 3), free
or united
at base, topped by feathery stigmas, exserted from sides or apex of floret. Fruit normally a dry indehiscent caryopsis with thin pericarp firmly adherent
to seed, pericarp rarely free, fleshy in some bamboos; embryo small or large; hilum
punctate
to linear
.
About 700 genera and 11,000 species: widely distributed in all regions of the world.[1]
Physical Description
Species Lolium temulentum
Annual
. Culms
tufted
, erect
or decumbent
, slender to moderately robust
,
20-120 cm tall, 3-5-noded. Leaf blades
flat, thin, 10-25 cm ×
4-10 mm, smooth
or scabridulous
on abaxial
surface, margins
scabrid
,
young blades rolled; auricles
present or absent; ligule 0.5-2.5 mm,
obtuse
to truncate
. Raceme
stiff, straight, 10-30 cm; rachis thick,
smooth or scabridulous, spikelets
about their own length
apart. Spikelets
turgid
, 0.8-2.5 cm, florets
4-10, rachilla internodes 1-1.5 mm, smooth,
glabrous
; glume
linear-oblong, rigid
, as long as spikelet, often
exceeding florets, 5-9-veined, margins narrowly membranous, apex
obtuse; lemmas elliptic
to ovate
, turgid at maturity, 5.2-8.5 mm,
apex obtuse; awn
usually present, stiff, scabrid; palea ciliolate
along keels. Caryopsis very plump, length 2-3 times width
, 4-7 mm.
Fl.
and fr. May-Aug. 2n = 14. [source]
Awnless or weakly awned
variants
can be distinguished at varietal
rank. [source]
Habit: Graminoid
Habitat
Fields
of cereals, introduced
.[2].
Typically found at an altitude of 0 to 4,653 meters (0 to 15,266 feet).[3]
Biology
Reproduction
Duration: Annual
Taxonomy
- Domain:
Eukaryota
(
)
- Whittaker & Margulis,1978
- eukaryotes
- Kingdom:
Plantae
(
)
- Haeckel, 1866
- Plants
- Subkingdom:
Viridaeplantae
(
)
- Cavalier-Smith, 1981
- Phylum:
Tracheophyta
(
)
- Sinnott, 1935 Ex Cavalier-Smith, 1998
- Vascular Plants
- Subphylum:
Euphyllophytina
(
)
- Infraphylum:
Radiatopses
(
)
- Kenrick & Crane, 1997
- Class:
Liliopsida
(
)
- Scopoli, 1760
- Subclass:
Commelinidae
(
)
- Takhtajan, 1967
- Superorder:
Poanae
(
)
- (Small, 1903) Takhtajan, 1997 Ex Reveal & Doweld, 1999
- Superorder:
Poanae
(
- Subclass:
Commelinidae
(
- Class:
Liliopsida
(
- Infraphylum:
Radiatopses
(
- Subphylum:
Euphyllophytina
(
- Phylum:
Tracheophyta
(
- Subkingdom:
Viridaeplantae
(
- Kingdom:
Plantae
(
Unambiguous Synonyms
- Bromus temulentus (L.) Bernh.
- Craepalia temulenta (L.) Schrank
- Lolium arvense With.
- Lolium temulentum var. arvense (With.) Lilja
- Lolium temulentum var. leptochaeton A. Braun
- Lolium temulentum var. macrochaeton A. Braun
Notes
Name
Status: Accepted Name
. Latest taxonomic
scrutiny: 15-Mar-2000
Place of publication
: Sp.
pl. 1:83. 1753
Name verified on 08-May-1992 by ARS Systematic Botanists. Last updated: 20-Nov-2007
Similar Species
Members of the genus Lolium
ZipcodeZoo has pages for 132 species, subspecies, varieties, forms, and cultivars in this genus. Here are just 100 of them:
L. aechicum · L. aegyptiacum · L. agreste · L. album · L. ambiguum · L. annuum · L. appenninum · L. arenarium · L. aristatum · L. arundinaceum (Lolium Arundinaceum) · L. arvense · L. aschersoniana · L. asperum · L. berteronianum · L. boucheanum · L. brasilianum · L. bromoides · L. canadense · L. canariense (Canary Islands Ryegrass) · L. cechicum · L. coelorachis · L. complanatum · L. compositum · L. compressum · L. crassiculme · L. cristatum · L. cuneatum · L. cylindricum · L. decipiens · L. distachyum · L. dorei · L. edwardii · L. elegans · L. elongatum · L. felix · L. festuca · L. festucaceum · L. festucoides · L. flagellare · L. gaudini · L. giganteum (Giant Fescue) · L. glumosum · L. gmelini · L. gracile · L. grandispicum · L. gussonei · L. halleri · L. hubbardii · L. humile · L. husnoti · L. hybrid · L. hybridum (Short-Rotation Ryegrass) · L. 'Hymer' · L. infelix · L. italicum · L. italicum var. muticum · L. jechelianum · L. latum · L. lepturoide · L. lepturoides · L. lesdaini · L. linicola · L. linicolum · L. loliaceum · L. longiglume · L. lowei · L. lucidum · L. macilentum · L. marschallii · L. maximum · L. mazzettianum · L. multiflorum · L. multiflorum italicum · L. multiflorus · L. oechicum · L. osiridis · L. parabolicae · L. parenne · L. perene · L. perenne (Perennial Rye Grass) · L. perennem · L. perenne aristatum · L. perenne cristatum · L. perenne 'Literra' · L. perenne multiflorum (Perennial Rye Grass) · L. perenne rigidum · L. perenne perenne · L. perenne var. brasilianum · L. perenne var. italicum · L. perenne var. perenne · L. perenne x · L. persicum (Persian Rye Grass) · L. phoenice · L. phoenix · L. pratense (Meadow Ryegrass) · L. pratensis · L. remotum · L. remotum schrank · L. repens · L. rigidium
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Further Reading
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Notes
Contributors
- Bisby, F.A., Y.R. Roskov, M.A. Ruggiero, T.M. Orrell, L.E. Paglinawan, P.W. Brewer, N. Bailly, J. van Hertum, eds (2007). Species 2000 & ITIS Catalogue of Life: 2007 Annual Checklist. Species 2000: Reading, U.K.
- Global Biodiversity Information Facility. Accessed November 15, 2007. http://www.gbif.org Mediated distribution data from 30 providers.
- "Lolium temulentum". in Flora of China Vol. 22 Page 243. Published by Missouri Botanical Garden Press. Online at EFloras.org.
- USDA, ARS, National Genetic Resources Program. Germplasm Resources Information Network - (GRIN) [Online Database]. National Germplasm Resources Laboratory, Beltsville, Maryland. URL (April 29, 2008)
Data Sources
Accessed through GBIF Data Portal November 15, 2007:
- Biologiezentrum der Oberoesterreichischen Landesmuseen, Biologiezentrum Linz
- Bundesamt für Naturschutz / Zentralstelle für Phytodiversität Deutschland, Bundesamt fuer Naturschutz / Zentralstelle fuer Phytodiversitaet Deutschland
- Conservatoire botanique national du Bassin parisien, Conservatoire botanique national du Bassin parisien
- International Plant Genetic Resources Institute(IPGRI), EURISCO
- International Plant Genetic Resources Institute(IPGRI), The System-wide Information Network for Genetic Resources
- Jyväskylä University Museum - The Section of Natural Sciences, Vascular plant collection of Jyvaskyla University Museum
- Missouri Botanical Garden, Missouri Botanical Garden
- Natural History Museum, University of Oslo, Vascular Plant Herbarium, Oslo
- Natural History Museum, University of Oslo, Vascular Plants, Field notes, Oslo
- The Swedish Museum of Natural History
- , Botany
- The Swedish Museum of Natural History
- , Herbarium of Oskarshamn
- The Swedish Museum of Natural History
- , Lund Botanical Museum
- Tiroler Landesmuseum Ferdinandeum, Tiroler Landesmuseum Ferdinandeum
- USDA PLANTS, USDA PLANTS Database
- University of Alabama Biodiversity and Systematics, Herbarium
Identifiers
- Biodiversity Heritage Library NamebankID: 2661079
- Catalogue of Life Accepted Name Code: ITS-40900
- Global Biodiversity Information Facility Taxonkey: 14013169
- Globally Unique Identifier: urn:lsid:ipni.org:names:407518-1
- GRIN Nomen Number: 22502
- Integrated Taxonomic Information System (ITIS) Taxonomic Serial Number (TSN): 40900
- Natural Heritage Network Species Identifier: PMPOA3U060
- U.S.D.A. Plant Symbol: LOTEM
- Zipcode Zoo Species Identifier: 47612
Footnotes
- Shou-liang Chen, De-Zhu Li, Guanghua Zhu, Zhenlan Wu, Sheng-lian Lu, Liang Liu, Zheng-ping Wang, Bi-xing Sun, Zheng-de Zhu, Nianhe Xia, Liang-zhi Jia, Zhenhua Guo, Wenli Chen, Xiang Chen, Yang Guangyao, Sylvia M. Phillips, Chris Stapleton, Robert J. Soreng, Susan G. Aiken, Nikolai N. Tzvelev, Paul M. Peterson, Stephen A. Renvoize, Marina V. Olonova & Klaus Ammann "Poaceae". in Flora of China Vol. 22. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org. [back]
- "Lolium temulentum". in Flora of China Vol. 22 Page 243. Missouri Botanical Garden Press. Online at EFloras.org. [back]
- Mean = 177.910 meters (583.694 feet), Standard Deviation = 298.050 based on 2,262 observations. Altitude information for each observation from British Oceanographic Data Centre. [back]
