Overview
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Threatened |
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Description
Family Urticaceae
Herbs, subshrubs
, or shrubs
, rarely trees
, very rarely climbing
, stems often fibrous
, sometimes succulent. sometimes armed
with stinging
hairs
; epidermal cells
of leaves, sometimes stems, perianths mostly with prominent
cystoliths
punctiform
to linear
; Leaves alternate or opposite, stipules present, rarely absent; leaf blade
simple
. Inflorescences cymose
, paniculate
, racemose, spicate
, or cluster-capitate, usually formed from glomerules
, sometimes crowded on common enlarged cuplike or discoid
receptacle, rarely reduced into a single flower. Flowers unisexual
(plants
monoecious or dioecious), rarely bisexual
in partial flowers; actinomorphic
, very small, (1-) 4- or 5-merous, rarely perianth absent in female flowers. Calyx absent. Perianth lobes
imbricate or valvate
. Male flowers: stamens as many as and opposite to perianth lobes, filaments
inflexed
in bud; anthers
2-locular, opening lengthwise, rudimentary
ovary often present. Female flowers: perianth lobes free
or connate
, usually enlarged in fruit and persistent
, occasionally absent; staminodes scarious
, opposite to the perianth lobes, or absent. Ovary rudimentary in male flowers, sessile or shortly stipitate
, free or adnate
to the perianth; 1-locular, ovule solitary, erect
from the base
; style
simple, or absent; stigma diverse
, capitate, penicillate-capitate (brushlike), subulate
, filiform
, ligulate
, or peltate. Fruit usually a dry achene, sometimes a fleshy
drupe, often enclosed by the persistent perianth. Seed solitary, endosperm usually present; embryo straight; cotyledons ovate
elliptical
or orbicular
.
About 47 genera and 1300 species: most numerous
in wet tropical regions
, extending into temperate regions
; 25 genera and 341 species (163 endemic, one introduced
) in China.
Plants in this family
have numerous uses. The stem fiber of some genera and species is of high quality and used to make cloth, fishing nets
, and ropes and for some industrial materials
. In central and southern China,
Boehmeria nivea is widely cultivated for ramie fiber and Girardinia diversifolia subsp.
triloba is widely cultivated for red huo ma fiber. Boiled young shoots
of Girardinia, Laportea, and Urtica are eaten as vegetables. Some species are used in local Chinese medicine. Pellionia repens, Pilea cadierei, P. microphylla, and P. peperomioides, among other species, are widely cultivated as ornamentals
in China and elsewhere. Some genera, such as Elatostema, Pellionia, and Pilea, occur frequently in shady, moist habitats
of subtropical
forests
and become dominant elements
of the forest floor vegetation. Plants of the first five genera belong to tribe
Urticeae, which is usually characterized by the distinctive stinging hairs.[1]
Genus Pilea
Herbs, shrubs
, or subshrubs
, annual
or perennial
, glabrous
. Stems simple
or branched, erect
, ascending
, or repent
. Leaves opposite; stipules present. Leaf blades
paired
, equal or unequal, ovate
, margins
dentate
or entire
; cystoliths
linear
, ± conspicuous
. Inflorescences axillary
, compact
to lax
cymes. Flowers unisexual
, staminate
and pistillate
flowers in same cyme; bracts deltate to linear. Staminate flowers
: tepals 4; stamens 4; pistillode
conic. Pistillate flowers: tepals 3, equal or sometimes 1 tepal enlarged and hoodlike; staminodes 3, opposite tepals, under tension and ejecting mature
achene; style
and tufted
stigma deciduous. Achenes sessile, laterally compressed
, ovoid
to teardrop-shaped, free
from perianth at maturity, partly covered by hoodlike tepal. x
= 12, 13.
Species ca.
400: mostly tropical
and subtropical
regions worldwide except Australia and New Zealand.
Pilea should be further revised.[2]
Habitat
Ecology:
List of Habitats
:1.9Forest - Subtropical/Tropical Moist Montane
Taxonomy
- Domain:
Eukaryota
(
)
- Whittaker & Margulis,1978
- eukaryotes
- Kingdom:
Plantae
(
)
- Haeckel, 1866
- Plants
- Subkingdom:
Viridaeplantae
(
)
- Cavalier-Smith, 1981
- Phylum:
Tracheophyta
(
)
- Sinnott, 1935 Ex Cavalier-Smith, 1998
- Vascular Plants
- Subphylum:
Euphyllophytina
(
)
- Infraphylum:
Radiatopses
(
)
- Kenrick & Crane, 1997
- Class:
Magnoliopsida
(
)
- Brongniart, 1843
- Dicotyledons
- Subclass:
Dilleniidae
(
)
- Takhtajan, 1967
- Superorder:
Urticanae
(
)
- Takhtajan Ex Reveal, 1992
- Order:
Urticales
(
)
- Dumortier, 1829
- Family:
Urticaceae
(
)
- Durande, 1782, Nom. Cons.
- Nettle Family
- Tribe:
Lecantheae
(
)
- Genus:
Pilea
(
)
- Lindley, Coll. Bot. plate 4 and text on facing page. 1821.
- [Latin pileus, felt cap, because of the calyx covering the achene]
- Specific epithet:
jamesonia
- Wedd.
- Botanical name: - Pilea jamesonia Wedd.
- Specific epithet:
jamesonia
- Wedd.
- Genus:
Pilea
(
- Tribe:
Lecantheae
(
- Family:
Urticaceae
(
- Order:
Urticales
(
- Superorder:
Urticanae
(
- Subclass:
Dilleniidae
(
- Class:
Magnoliopsida
(
- Infraphylum:
Radiatopses
(
- Subphylum:
Euphyllophytina
(
- Phylum:
Tracheophyta
(
- Subkingdom:
Viridaeplantae
(
- Kingdom:
Plantae
(
Similar Species
Members of the genus Pilea
There are approximately 725 species in this genus. Here are just 100 of them:
P. abbreviata · P. acanthospermoides · P. achanthospermoides · P. acuminata · P. acunae · P. adamsiana · P. aenea · P. affinis · P. affinis var. havanensis · P. alongensis · P. alpestris · P. alpina · P. alsinifolia · P. alta · P. alternifolia · P. amamiana · P. ambecarpa · P. amplistipulata · P. andersonii · P. 'Anette' · P. angolensis · P. angolensis angolensis · P. angolensis christiaensenii · P. angulata · P. angulata latiuscula · P. angulata petiolaris · P. angulata subsp. latiuscula · P. angulata subsp. petiolaris · P. angustata · P. angustifolia · P. anisophylla · P. anivoranensis · P. anomala · P. antioquensis · P. antiqouensis · P. aparadensis · P. aphrophila · P. apiculata · P. apoensis · P. appendicilata · P. appendiculata · P. approximata · P. approximata var. incisoserrata · P. aquarum · P. aquarum acutidentata · P. aquarum subsp. acutidentata · P. aquarum subsp. brevicornuta · P. argentea · P. arguta · P. aripoensis · P. articulata · P. astrogramma · P. atroviridis · P. attenuata · P. auricularis · P. auriculata · P. balansae · P. balfourii · P. baltenweckii · P. bambuseti · P. bambusifolia · P. barahonensis · P. barbiflora · P. basicordata · P. basilensis · P. bassleriana · P. baurii (Galapagos Pilea) · P. baviensis · P. beguinotii · P. beguinotti · P. bemarivensis · P. benguetensis · P. betulaefolia var. poitaei · P. betulifolia · P. betulifolia var. lineolata · P. bicolor · P. bisepala · P. bissei · P. blinii · P. boehmerioides · P. boiviniana · P. boniana · P. borbonica · P. botterii · P. brachyclada · P. brachypila · P. bracteosa · P. brassii · P. brevicornuta · P. brevistipula · P. brittoniae · P. bronxensis · P. buchenavii · P. buchtienii · P. bullata · P. cacuminum · P. cadetii · P. cadierei (Watermelon Plant) · P. cadierei 'Minima' · P. cadierei 'Minima Nana'
More Info
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- Search using Specialized Databases: GenBank | Medline | Scirus | CISTI/CAL | Agricola Periodicals | Agricola Books
Further Reading
- Chen Chiajui & Wang Wentsai. 1995. Urticaceae. In: Wang Wentsai & Chen Chiajui, eds., Fl. Reipubl. Popularis Sin. 23(2): 1404.
- Chen, C. J. 1982. A monograph of Pilea (Urticaceae) in China. Bull. Bot. Res., Harbin 2: 1-132.
- Fernald, M. L. 1936. Pilea in eastern North America. Contr. Gray Herb. 113: 169-170.
- Hermann, F. J. 1940. The geographic distribution of Pilea fontana. Torreya 40: 118-120.
Notes
Contributors
- Brands, S.J. (comp.) 1989-2007. Systema Naturae 2000. The Taxonomicon. Universal Taxonomic Services, Amsterdam, The Netherlands. Accessed March 24, 2007.
- Valenzuela, J.C., Rojas, W. & Pitman, N. 2004. In IUCN 2008. 2008 IUCN Red List of Threatened Species. IUCNRedList.org. Downloaded July 19, 2008.
Identifiers
- Biodiversity Heritage Library NamebankID: 5989688
- IUCN ID: 46241
- Zipcode Zoo Species Identifier: 1029194
Footnotes
- Jiarui Chen, Prof. Qi Lin, Ib Friis, C. Melanie Wilmot-Dear & Alex K. Monro "Urticaceae". in Flora of China Vol. 5 Page 76. Published by Science Press (Beijing) and Missouri Botanical Garden Press. Online at EFloras.org. [back]
- "Pilea". in Flora of North America Vol. 3. Oxford University Press. Online at EFloras.org. [back]
