Plants usually woody, or herbs, sometimes lacking chlorophyll. Leaves spiral, sometimes decussate, margin often toothed. Inflorescence racemose; bracteoles paired, basal. Flowers (4 or) 5-merous. Calyx imbricate. Corolla connate, imbricate. Stamens 10, sometimes with spurs or awns, dehiscing by pores, pollen in tetrads, rarely single. Ovary superior or inferior, placentation axile, rarely parietal, often many ovules per locule. Style ± as long as corolla, slender. Fruit a capsule or berry, rarely a drupe; calyx persistent.
About 125 genera and 4000 species: widely distributed in temperate and subarctic regions, also at high elevations in tropical regions; 22 genera and 826 species (524 endemic) in China.
The Monotropoideae are here included in the Ericaceae; previously in FRPS (56: 157 216. 1990), they were treated as the Pyrolaceae. Chiogenes, recorded from China in FRPS (57(3) : 69 71. 1991), is here included in Gaultheria. Over the last half century, the Empetraceae have usually been separated from, but closely associated with, the Ericaceae. In their ecology, leaf morphology and insertion, rusts, embryology, stamen anatomy, etc., they largely agree with that family. Molecular data place Empetrum and its relatives firmly within the Ericaceae, and in particular within the subfamily Ericoideae, in agreement with phytochemical and palynological data, and there they are best recognized as a separate tribe. There are distinctive features of the Empetreae that were responsible for their past familial status, e.g., reduced perianth with separate members, low ovule number, enlarged stigmas, etc. However, these are likely to be derived features associated with wind pollination. See Kron et al. (Bot. Rev. 68: 335 423. 2002) and the recent treatment of the Ericaceae by Stevens et al. (in Kubitzki, Fam. Gen. Vasc. Pl. 6: 145 194. 2004) .
Several genera and many species are ornamentals. Some fruits of Vaccinium in N China are sweet and edible, but of no particular value to humans. Some species of Chamaedaphne, Craibiodendron, Leucothoë, Lyonia, Pieris, and Rhododendron contain more or less toxic diterpenes, which are harmful to humans or domestic animals.[1]
Shrubs or small trees, terrestrial or epiphytic. Leaves evergreen or deciduous, rarely pseudoverticillate, petiolate, margin entire or serrate. Inflorescences terminal or axillary, racemose, fasciculate, or solitary flowers. Bracts and bracteoles persistent or caducous, the latter small, basal, rarely apical; pedicel expanded towards apex or not, articulate, rarely continuous. Flowers 5-, rarely 4-merous. Calyx limb lobed or toothed. Corolla urceolate, campanulate, or tubular, lobed or toothed, usually to base of limb; lobes erect or recurved, mostly shorter than tube, rarely longer or petals apparently free. Stamens rarely 4, usually included; anthers with 2 spurs at anther-filament junction or not; thecae with tubules opening by a terminal pore or introrse slit. Disk annular. Ovary inferior, usually 8- or 10-pseudoloculed by false partitions; locule with many ovules; stigma inconspicuous, truncate. Fruit a several seeded globose berry. Seeds ovoid, small, testa hard or mucilaginous.
About 450 species: throughout the N hemisphere, also in the mountains of tropical Asia and Central and South America, a few species in Africa and Madagascar; 92 species (51 endemic) in China.
The limits of Vaccinium are very uncertain. Recent molecular work, supported by morphology and anatomy, suggests that it is likely that the majority of Chinese species, i.e., those with 10-pseudoloculed ovaries, form part of the same lineage as Agapetes.[2]
Habit: Deciduous.
Flowers: Bloom Period: January, February.
Culture: Space 6-8' apart.
Soil: Minimum pH: 4.6 • Maximum pH: 5.5
Sunlight: Sun Exposure: Full Sun.
Temperature: Cold Hardiness: 7b, 8a, 8b, 9a, 9b. (map)
There are approximately 977 species, subspecies, varieties, forms, and cultivars in this genus. Here are just 100 of them: V. uliginosum gaultherioides · V. uliginosum imberbe · V. uliginosum occidentale · V. uliginosum pubescens · V. sphyrospermoides · V. 'Berkeley' · V. 'Bluejay' · V. 'Blueray' · V. 'Chippewa' · V. 'Cinderella' · V. 'Duke' · V. 'Earliblue' · V. 'Groover' · V. 'McMinn' · V. 'Misty' · V.'NC 2282' (Blueberry) · V. 'Nimo Pink' · V. 'Northblue' (Half-High Highbush Blueberry) · V. 'Northcountry' (Half-High Highbush Blueberry) · V. 'Northland' (Half-High Highbush Blueberry) · V. 'Northsky' (Half-High Highbush Blueberry) · V. 'Ozarkblue' · V. 'Patriot' · V. 'Spartan' · V. 'Sunrise' · V. 'Sunshine Blue' · V. 'Top Hat' (Dwarf Blueberry) · V. 'Tophat' · V. absconditum · V. acosta · V. acrobracteatum · V. acuminatissimum · V. acuminatum · V. acutissimum · V. adenandrum · V. adenanthum · V. adenochaetum · V. adenotrichum · V. adenurum · V. affine · V. africanum · V. agathosmoides · V. agusanense · V. aitapense · V. alainii · V. alaskaense · V. alaskaensis · V. alaskensis · V. alaternoides · V. alaternoides Kunth var. parvifolium Benth. · V. alaternoides var. parvifolium · V. alatum · V. albicans · V. albicans var. albicans · V. albidens · V. albiflorum · V. album · V. almedae · V. alnifolium · V. altiterrae · V. alto-montanum · V. alvarezii · V. amamianum · V. ambiualens (Stout Blacksmelt) · V. ambivalens · V. amblyandrum · V. ambyandrum · V. amicorum · V. amoemum · V. amphoterum · V. amplifolium · V. ampullaceum · V. andersonii · V. andringitrense · V. anfractum · V. angiense · V. angulatum · V. angulosum · V. angustifolium (Lowbush Blueberry) · V. angustifolium 'Burgundy' · V. angustifolium 'Claret' · V. angustifolium 'Cumberland' · V. angustifolium 'Jonesboro' · V. angustifolium Aiton var. hypolasium Fernald · V. angustifolium Aiton var. laevifolium House · V. angustifolium f. chiococcum · V. angustifolium f. leucoarpum · V. angustifolium f. leucocarpum · V. angustifolium f. nigrum · V. angustifolium Newfoundland form · V. angustifolium var. glaucescens · V. angustifolium var. integrifolium · V. angustilimbum · V. anthonyi · V. antipodum · V. apiculatum · V. apoanum · V. apricum · V. arborescens · V. arborescens var. glaucescens
There are approximately 16,146 species, subspecies, varieties, forms, and cultivars in this genus. Here are just 100 of them: I. yebrudii edgecombii · I. agatha · I. alto-barbata · I. aureonympha · I. autosyndetica · I. caesia · I. intercedens · I. neglecta · I. nelsonii · I. neumayeri · I. orientosa · I. robusta · I. rotschildi · I. sancti-cyri · I. setosothungbergii · I. shelfordensis · I. tenosa · I. 'A L'Orange' (A Lorange Tall Bearded Iris) · I. 'Aachen Elf' (Miniature Tall Bearded) · I. 'Aachen' (Tall Bearded Iris) · I. 'Aakukui' (Aakukui Tall Bearded Iris) · I. 'Aardvark Antickz' (Intermediate Bearded Iris) · I. 'Aardvark Lark' (Tall Bearded Iris) · I. 'Aaron's Angel' (Tall Bearded Iris) · I. 'Aaron's Bluebells' (Tall Bearded Iris) · I. 'Aaron's Bonus' (Aarons Bonus Tall Bearded Iris) · I. 'Aaron's Dagger' (Aarons Dagger Tall Bearded Iris) · I. 'Aaron's Dream' (Aarons Dream Tall Bearded Iris) · I. 'Aaron's Rod' (Aarons Rod Tall Bearded Iris) · I. 'Abadiyat' (Tall Bearded Iris) · I. 'Abba Alias Abba' (Standard Dwarf Bearded Iris) · I. 'Abbey Chant' (Abbey Chant Intermediate Bearded Iri) · I. 'Abbey Road' (Abbey Road Tall Bearded Iris) · I. 'Abbondanza' (Tall Bearded Iris) · I. 'Abby And Me' (Tall Bearded Iris) · I. 'Abby Grace' (Tall Bearded Iris) · I. 'Abdera' · I. 'Abdul Unknown' (Tall Bearded Iris) · I. 'Abiding Joy' (Abiding Joy Tall Bearded Iris) · I. 'Abilene' (Border Bearded Iris) · I. 'Abiqua Falls' (Tall Bearded Iris) · I. 'Ablaze' · I. 'About Last Night' (Tall Bearded Iris) · I. 'About Time' (Tall Bearded Iris) · I. 'About Town' (Tall Bearded Iris) · I. 'Above Board' (Tall Bearded Iris) · I. 'Above the Clouds' (Above the Clouds Tall Bearded Iris) · I. 'Abracadabra' (Abracadabra Standard Dwarf Bearded Iri) · I. 'Abridged Version' (Abridged Version Miniature Tall Bearded Iri) · I. 'Abstract Art' (Abstract Art Tall Bearded Iris) · I. 'Abstraction' (Tall Bearded Iris) · I. 'Acacia Rhumba' (Louisiana Iris) · I. 'Acadian Miss' (Acadian Miss Louisiana Iri) · I. 'Acadian Rose' (Louisiana Iris) · I. 'Acapulco Gold' (Acapulco Gold Tall Bearded Iris) · I. 'Acapulco Sunset' (Acapulco Sunset Tall Bearded Iris) · I. 'Acapulco' (Tall Bearded Iris) · I. 'Acceleration' (Standard Dwarf Bearded Iris) · I. 'Accent' (Tall Bearded Iris) · I. 'Access Code' (Tall Bearded Iris) · I. 'Accidents Happen' · I. 'Acclaim' · I. 'Acclamation' (Tall Bearded Iris) · I. 'Ace of Clubs' (Ace of Clubs Standard Dwarf Bearded Iri) · I. 'Ace of Diamonds' (Tall Bearded Iris) · I. 'Ace Royale' (Tall Bearded Iris) · I. 'Ace' (Miniature Tall Bearded Iris) · I. 'Acey Deucey' (Acey Deucey Standard Dwarf Bearded Iri) · I. 'Acid Print' (Standard Dwarf Bearded Iris) · I. 'Acolyte' · I. 'Acoma' (Acoma Tall Bearded Iris) · I. 'Aconite Elf' (Miniature Dwarf Bearded Iris) · I. 'Acstede Nine Hundred' · I. 'Act III' · I. 'Act of Kindness' (Act of Kindness Tall Bearded Iris) · I. 'Act Three' (Tall Bearded Iris) · I. 'Action Adventure' (Intermediate Bearded Iris) · I. 'Action Front' (Action Front Tall Bearded Iris) · I. 'Actress' (Actress Tall Bearded Iris) · I. 'Ad Astra' · I. 'Ad Lib' (Tall Bearded Iris) · I. 'Ad Vantage' (Tall Bearded Iris) · I. 'Ada Marie' (Tall Bearded Iris) · I. 'Ada Morgan' (Louisiana Iris) · I. 'Adam's Fall' (Tall Bearded Iris) · I. 'Adamsblack' (Tall Bearded Iris) · I. 'Add It Up' (Add It Up Standard Dwarf Bearded Iri) · I. 'Added Praise' (Tall Bearded Iris) · I. 'Added Value' (Tall Bearded Iris) · I. 'Address Unknown' (Tall Bearded Iris) · I. 'Aden' (Tall Bearded Iris) · I. 'Admiral Blue' (Tall Bearded Iris) · I. 'Admiral's Choice' (Tall Bearded Iris) · I. 'Adobe Rose' (Adobe Rose Tall Bearded Iris) · I. 'Adobe Sunset' (Adobe Sunset Spuria Iri) · I. 'Adopt Me' (Intermediate Bearded Iris) · I. 'Adorable Diva' (Tall Bearded Iris) · I. 'Adoregon' (Tall Bearded Iris) · I. 'Adoring Glances' (Standard Dwarf Bearded Iris) · I. 'Adriaan Backer'
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