Gesneriaceae
noun
Herbs, shrubs, or rarely trees. Leaves opposite or rarely alternate, whorled or basal, rosette forming; exstipulate; usually simple, rarely shallowly to deeply lobed, pinnately or rarely palmately veined. Inflorescences usually cymes, rarely racemes, axillary, often near apex and appearing terminal; usually pedunculate. Flowers perfect, zygomorphic, seldom actinomorphic. Calyx actinomorphic, rarely zygomorphic; usually (4 or) 5-divided. Corolla gamopetalous, zygomorphic, rarely actinomorphic; usually 2-lipped. Fertile stamens 2 or 4, then often didynamous, rarely 5, epipetalous; anthers free or coherent, thecae 2, parallel, divergent, or divaricate; staminodes 1-3 or absent. Disc ringlike to cupular, rarely absent. Ovary superior in all Old World taxa [half inferior, or inferior], 1-loculed; gynophore seldom present; placentas (1 or) 2, parietal, rarely 2-loculed, placenta 1 per locule and axile; ovules numerous, anatropous. Style 1;
stigmas 1 or 2. Fruit usually capsular, loculicidal, septicidal, or circumscissile, rarely a berry, indehiscent. Seeds numerous, fusiform to ellipsoid or ovoid, minute, sometimes with appendages at 1 or both ends, with or without endosperm; embryo straight, cotyledons equal or unequal after germination. About 133 genera and 3000 species: Africa, Central and South America, E and S Asia, S Europe, Oceania; 56 genera (25 endemic) and 442 species (354 endemic) in China. A few foreign well-known ornamental species are cultivated in China, including the florist's gloxinia, Sinningia speciosa (Loddiges) Hiern, and African violet, Saintpaulia ionantha Wendland. The two ovary carpels may each produce a stigma; these stigmas are fused into a single structure. Some students of Gesneriaceae have considered the stigma to be single and either simple (capitate) or 2-lobed, whereas others consider each of the two stigmas as units. We have maintained the latter usage, but a family-wide investigation of stigma development is needed. The distinction can be blurred, however, because the stigmas may be completely fused into one with a capitate apex (as in Didymocarpus) or one of the two carpels or stigmas may be aborted resulting in a single stigma that may or may not be 2-lobed.